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Salix caprea
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{{Short description|Species of tree}} {{Redirect|Goat Willow|the song by Hawkwind|Hall of the Mountain Grill}} {{Speciesbox |status = LC |status_system = IUCN3.1 |status_ref = <ref name=iucn>{{Cite iucn |title=''Salix caprea'' |author=Barstow, M., Matchutadze, I. & Harvey-Brown, Y. |name-list-style=amp |page= e.T19620273A19621176 |date=2018 |doi=10.2305/IUCN.UK.2018-1.RLTS.T19620273A19621176.en |access-date=11 April 2024}}</ref> |image = Waterwilg (DSC 2539).jpg |image_caption = Goat willow male catkins |genus = Salix |species = caprea |authority = [[Carl Linnaeus|L.]] |range_map = Salix caprea range.svg |range_map_caption = Distribution map |synonyms_ref=<ref>{{cite web |url=https://powo.science.kew.org/taxon/urn:lsid:ipni.org:names:303599-2 |title=''Salix caprea L.'' |author=<!--Not stated--> |date=2017 |website=Plants of the World Online |publisher=Board of Trustees of the Royal Botanic Gardens, Kew |access-date=7 September 2020 }}</ref> |synonyms={{collapsible list| *''Capraea vulgaris'' <small>Opiz</small> *''Nectopix caprea'' <small>(L.) Raf.</small> *''Salix aurigerana'' <small>Lapeyr.</small> *''Salix bakko'' <small>Kimura</small> *''Salix caprea'' f. ''elongata'' <small>(Nakai) Kitag.</small> *''Salix caprea'' var. ''ishidoyana'' <small>(Nakai) M.Kim</small> *''Salix caprea'' var. ''lanatifolia'' <small>Björnstr.</small> *''Salix caprea'' var. ''pendula'' <small>T.Lang</small> *''Salix coaetanea'' <small>(Hartm.) Flod.</small> *''Salix hallaisanensis'' <small>H.Lév.</small> *''Salix hultenii'' <small>Flod.</small> *''Salix ishidoyana'' <small>Nakai</small> *''Salix lanata'' <small>Vill.</small> *''Salix sphacelata'' <small>Sm.</small> *''Salix tomentosa'' var. ''androgyna'' <small>Ser.</small> *''Salix tomentosa'' var. ''macrophylla'' <small>Ser.</small> *''Salix tomentosa'' var. ''rotundifolia'' <small>Ser.</small> *''Salix tomentosa'' var. ''tenuifolia'' <small>Ser.</small> *''Salix tomentosa'' var. ''ternata'' <small>Ser.</small> }} }} '''''Salix caprea''''', known as '''goat willow''', '''[[pussy willow]]''' or '''great sallow''', is a common species of [[willow]] native to [[Europe]] and western and central [[Asia]].<ref name=meikle>Meikle, R. D. (1984). ''Willows and Poplars of Great Britain and Ireland''. BSBI Handbook 4. {{ISBN|0-901158-07-0}}.</ref> ==Description== It is a [[deciduous]] [[shrub]] or small [[tree]], reaching a height of {{convert|8|-|10|m|ft|abbr=on}}, rarely to 13 m. The [[leaf|leaves]] are 3–12 cm long and from 2–8 cm wide, broader than most other willows. The [[flower]]s are soft silky, and silvery 3-7-cm-long [[catkin]]s are produced in early spring before the new leaves appear; the male and female catkins are on different plants ([[plant sexuality|dioecious]]). The male catkins mature yellow at [[pollen]] release, the female catkins mature pale green. {{Multiple image | align = left | total_width = 300 | image1 = Salix caprea female TK 3.jpg | alt1 = female and male catkins of ''Salix caprea'' | caption1 = Female catkins | image2 = Salix caprea male TK 4.jpg | caption2 = Male catkins }} The [[fruit]] is a small [[capsule (fruit)|capsule]] 5–10 mm long containing numerous minute [[seed]]s embedded in fine, cottony hairs. The seeds are very small (about 0.2 mm) with the fine hairs aiding dispersal; they require bare [[soil]] to [[Germination|germinate]].<ref name=meikle/><ref name=rushforth>Rushforth, K. (1999). ''Trees of Britain and Europe''. Collins. {{ISBN|0-00-220013-9}}.</ref> The two [[variety (biology)|varieties]] are:<ref name=meikle/> *''S. c.'' var. ''caprea'' - lowland regions throughout the range, leaves thinly hairy above, densely hairy below, 5–12 cm long, stipules persistent until autumn *''S. c.'' var. ''sphacelata'' (Sm.) Wahlenb. (syn. ''S. caprea'' var. ''coaetanea'' Hartm.; ''S. coaetanea'' (Hartm.) Floderus) - high altitudes in the mountains of central and northern Europe (Alps, Carpathians, Scotland, Scandinavia), leaves densely silky-hairy on both sides, 3–7 cm long, stipules early deciduous. In western Norway there are trees of S. c. var 'Sphacelata' that are 18–20 meters tall and have mostly single trunks. ==Name== The Latin [[binomial nomenclature|specific epithet]] ''caprea'' means "goat".<ref name=RHSLG>{{cite book | last=Harrison |first=Lorraine | title=RHS Latin for Gardeners | year=2012 |publisher=Mitchell Beazley | location=United Kingdom | isbn=978-1845337315 }}</ref> This, and the common name goat willow, probably derive from the first known illustration of the species in [[Hieronymus Bock]]'s 1546 ''[[Herbal]]'', where the plant is shown being browsed by a goat. The species was historically also widely used as a browse for goats, to which Bock's illustration may refer.<ref name=bean>Bean, W. J. (1980). ''Trees and Shrubs Hardy in the British Isles''. {{ISBN|0-7195-2428-8}}.</ref> ==Ecology== ''S. caprea'' occurs both in wet/damp environments, such as riverbanks and lake shores, and in drier sites, wherever bare soil becomes available due to ground disturbance.<ref name=meikle/> [[Hybrid (biology)|Hybrids]] with several other willow species are common, notably with ''[[Salix cinerea]]'' (''S. × reichardtii''), ''[[Salix aurita]]'' (''S. × multinervis''), ''[[Salix viminalis]]'' (''S. × smithiana''), and ''[[Salix purpurea]]'' (''S. × sordida''). Populations of ''S. caprea'' often show [[hybrid introgression]].<ref name=meikle/><ref name=rushforth/> Unlike almost all other willows, pure specimens do not take root readily from cuttings; if a willow resembling the species does root easily, it is probably a hybrid with another species of willow.<ref name=rushforth/> The leaves are used as a food resource by several species of Lepidoptera, and are also commonly eaten by browsing [[mammal]]s. Willows are very susceptible to gall inducers, and the midge ''[[Rhabdophaga rosaria]]'' forms the camellia gall on ''S. caprea''.<ref>[http://www.joensuu.fi/biologia/nyman/IOWgallinducers.htm Gall Inducers] {{webarchive |url=https://web.archive.org/web/20110611161957/http://www.joensuu.fi/biologia/nyman/IOWgallinducers.htm |date=June 11, 2011 }}</ref> ==Cultivation and uses== {{See also|Pussy willow}} [[File:Willow whistle.jpg|thumb|170px|A willow flute]] A small number of [[cultivar]]s have been selected for garden use. The most common is ''S. caprea'' 'Kilmarnock', discovered by [[James Smith (Scottish botanist)|James Smith]], with stiffly pendulous shoots forming a mop-head; it is a male clone. A similar female clone is ''S. caprea'' 'Weeping Sally'. As they do not form a leader, they are [[grafting|grafted]] on erect stems of other willows; the height of these cultivars is determined by the height at which the graft is made.<ref name=rushforth/> Plants can also be grown from greenwood cuttings, which make attractive creeping mounds. Hardwood cuttings are often difficult to root. Both [[tannin]] and [[salicin]] can be extracted from goat willow bark. The tree is not considered a good source of [[timber]], as its wood is both brittle and known to crackle violently if burned. As with the closely related ''[[Salix discolor]]'' (American pussy willow), it is also often grown for cut flowers. See [[Pussy willow]] for further cultural information, which applies to both species. In [[North Europe]] it has been fairly common to make [[willow flute]]s from goat willow cuttings. In [[Germany]], [[Hungary]], north of [[Slovakia]], [[Poland]], [[Ukraine]] and many parts of [[Finland]], the just-opened catkins are used like the olive branches on [[Palm Sunday]]. ==References== {{Reflist}} ==External links== {{Commons category}} {{Wikispecies}} * [http://www.euforgen.org/species/salix-caprea/ ''Salix caprea''] - genetic conservation units [[European Forest Genetic Resources Programme]] (EUFORGEN) * {{PFAF|Salix caprea}} * {{PFAF|Salix bakko}} {{Tannin source}} {{Taxonbar|from=Q159565}} [[Category:Salix|caprea]] [[Category:Flora of Ukraine]] [[Category:Plants described in 1753]] [[Category:Taxa named by Carl Linnaeus]]
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