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{{Short description|Small- to medium-sized perennial wood plant}} {{about|the plant|other uses}} {{Redirect|Bushes|the American political family|Bush family|other uses|Bush (disambiguation)}} {{More citations needed|date=October 2018}} [[File:Cytisus scoparius2.jpg|thumb|250px|A [[Cytisus scoparius|broom]] shrub in flower]] [[File:Sheringham Park 1.JPG|thumb|250px|A [[rhododendron]] shrubbery in [[Sheringham Park]]]] A '''shrub''' or '''bush''' is a small to medium-sized [[perennial]] [[woody plant]]. Unlike [[herbaceous plant]]s, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from [[tree]]s by their multiple [[Plant stem|stems]] and shorter [[height]], less than {{convert|6β10|m|ft|abbr=on}} tall.<ref name="LawrenceHawthorne2006">{{cite book | first1 = Anna | last1 = Lawrence | first2 = William | last2 = Hawthorne | title = Plant Identification: Creating User-friendly Field Guides for Biodiversity Management | url = https://books.google.com/books?id=rHEpVBM5-eIC&pg=PA138 | year = 2006 | publisher = Routledge | isbn = 978-1-84407-079-4 | pages = 138β }}</ref><ref name="Allaby2019">{{cite book |last1=Allaby |first1=Michael |title=A dictionary of plant sciences |date=2019 |publisher=Oxford Oxford University Press |isbn=9780198833338 |language=en|oclc=1097073225 }}</ref> Small shrubs, less than {{convert|2|m|ft|abbr=off}} tall are sometimes termed as [[subshrub]]s. Many [[botany|botanical groups]] have species that are shrubs, and others that are trees and herbaceous plants instead. Some define a shrub as less than {{Convert|6|m|ft|abbr=on}} and a tree as over 6 m. Others use {{Convert|10|m|ft|abbr=on}} as the cutoff point for classification.<ref name="Allaby2019" /> Many trees do not reach this mature height because of hostile, less than ideal growing conditions, and resemble shrub-sized plants. Others in such species have the potential to grow taller in ideal conditions. For longevity, most shrubs are classified between [[Perennial plant|perennials]] and trees. Some only last about five years in good conditions. Others, usually larger and more woody, live beyond 70. On average, they die after eight years.<ref>{{Cite web |date=2014-02-27 |title=Life Spans of Small Trees and Shrubs |url=https://temeculalandscapeconstruction.com/life-spans-of-small-trees-and-shrubs/ |access-date=2022-04-29 |website=McCabe's Landscape Construction |language=en-US}}</ref> [[Shrubland]] is the natural landscape dominated by various shrubs; there are many distinct types around the world, including [[fynbos]], [[maquis shrubland|maquis]], [[shrub-steppe]], [[shrub swamp]] and [[moorland]]. In gardens and parks, an area largely dedicated to shrubs (now somewhat less fashionable than a century ago) is called a [[shrubbery]], shrub border or shrub garden. There are many garden [[cultivar]]s of shrubs, bred for flowering, for example [[rhododendron]]s, and sometimes even leaf colour or shape. Compared to trees and herbaceous plants, a small number of shrubs have culinary usage.{{citation needed|date=April 2024}} Apart from the several [[berry]]-bearing species (using the culinary rather than botanical definition), few are eaten directly, and they are generally too small for much [[timber]] use unlike trees. Those that are used include several perfumed species such as [[Lavandula angustifolia|lavender]] and [[rose]], and a wide range of plants with medicinal uses. [[Camellia sinensis|Tea]] and [[Coffea|coffee]] are on the tree-shrub boundary;<ref>{{Cite web |last=Clayton |first=Liz |title=Is The Coffee Plant A Tree, Bush, Or Shrub? |url=https://sprudge.com/is-the-coffee-plant-a-tree-bush-or-shrub-181869.html |access-date=2022-04-29 |website=sprudge.com |date=20 October 2021 |language=en-US}}</ref> they are normally harvested from shrub-sized plants, but these would be large enough to become small trees if left to grow instead. ==Definition== Shrubs are perennial woody plants, and therefore have persistent woody [[Plant stem|stems]] above ground (compare with succulent stems of [[herbaceous plant]]s).<ref name="Allaby2019" /> Usually, shrubs are distinguished from trees by their height and multiple stems. Some shrubs are [[deciduous]] (e.g. [[Rhaphiolepis|hawthorn]]) and others [[evergreen]] (e.g. [[holly]]).<ref name="Allaby2019" /> [[ancient Greece|Ancient Greek]] philosopher [[Theophrastus]] divided the plant world into trees, shrubs and herbs.<ref name="Bremness1994">{{cite book |last1=Bremness |first1=Lesley |title=The complete book of herbs |publisher=Viking Studio Books |isbn=9780140238020 |page=8 |url=https://books.google.com/books?id=VqfnrH-W2kwC|year=1994 }}</ref> Small, low shrubs, generally less than {{Convert|2|m|ft|abbr=on}} tall, such as [[lavender]], [[Vinca|periwinkle]] and most small garden varieties of [[rose]], are often termed as [[subshrub]]s.<ref name="Fischer1990">{{cite book | first = Peggy | last = Fischer | title = Essential shrubs: the 100 best for design and cultivation | url = https://books.google.com/books?id=QuLtawHEsJMC | year = 1990 | publisher = Friedman/Fairfax Publishers | isbn = 978-1-56799-319-6 | pages = 9β | quote = ... Examples of subshrubs include candytuft, lavender, and rosemary. These broad definitions are ... }}</ref><ref>{{Cite web |date=2017-05-15 |title=What is a Subshrub? |url=https://worldoffloweringplants.com/subshrub/ |access-date=2022-04-29 |website=World of Flowering Plants |language=en-US}}</ref> Most definitions characterize shrubs as possessing multiple stems with no main trunk below.<ref name="Allaby2019" /> This is because the stems have branched below ground level. There are exceptions to this, with some shrubs having main trunks, but these tend to be very short and divide into multiple stems close to ground level without a reasonable length beforehand. Many trees can grow in multiple stemmed forms also while being tall enough to be trees, such as [[oak]] or [[ash (tree)|ash]].<ref name="Allaby2019" /> ==Use in gardens and parks== An area of [[Plant cultivation|cultivated]] shrubs in a [[park]] or a [[garden]] is known as a [[shrubbery]].<ref name="Whitefield2002">{{cite book | first = Patrick | last = Whitefield | title = How to Make a Forest Garden | url = https://books.google.com/books?id=3rd3e69BnC8C&pg=PA113 | year = 2002 | publisher = Permanent Publications | isbn = 978-1-85623-008-7 | pages = 113β }}</ref> When clipped as [[topiary]], suitable species or varieties of shrubs develop dense [[foliage]] and many small leafy [[branch]]es growing close together.<ref>{{cite book|url=https://books.google.com/books?id=a93TcRVsqLwC&q=topiary+pruning+dense+foliage&pg=PA100|title=Pruning for Flowers and Fruit|first=Jane|last=Varkulevicius|date=17 May 2010|publisher=Csiro Publishing|access-date=19 December 2017|via=Google Books|isbn=9780643101975}}</ref> Many shrubs respond well to renewal [[pruning]], in which hard cutting back to a "[[Living stump|stool]]", removes everything but vital parts of the plant, resulting in long new stems known as "canes".<ref>{{Cite web |date=2019-02-21 |title=Rejuvenation or Renewal Pruning to Restore Overgrown Shrubs |url=https://organicplantcarellc.com/rejuvenation-or-renewal-pruning-to-restore-overgrown-shrubs/ |access-date=2022-04-29 |website=Organic Plant Care LLC {{!}} Organic Lawn & Plant Health Service in Hunterdon, Morris, Somerset & Union Counties, NJ and Bucks County, PA |language=en-US}}</ref> Other shrubs respond better to selective pruning to dead or unhealthy, or otherwise unattractive parts to reveal their structure and character.<ref>{{Cite web |last=Turpin |first=Jason |date=2018-08-29 |title=What is Selective Tree and Shrub Pruning-How to Prune Correctly! |url=https://www.turpinlandscapedesign.com/what-is-selective-tree-and-shrub-pruning-landscape-experts-answer-your-questions/ |access-date=2022-04-29 |website=Turpin Landscape Design/Build |language=en-US}}</ref> Shrubs in common garden practice are generally considered [[flowering plant|broad-leaved plants]], though some smaller [[Pinophyta|conifers]] such as [[mountain pine]] and [[common juniper]] are also shrubby in structure. Species that grow into a shrubby habit may be either [[deciduous]] or [[evergreen]].<ref>{{cite book|url=https://books.google.com/books?id=vlDxSBGB0cYC&q=shrub+deciduous+or+evergreen|title=Popular Deciduous and Evergreen Trees and Shrubs|first=Franklin Reuben|last=Elliott|date=1 November 2008|publisher=Applewood Books|access-date=19 December 2017|via=Google Books|isbn=9781429012904}}</ref> == Botanical structure == {{See also|Shrubland}} [[File:Scrub brush vegetation in south TX IMG 6069.JPG|right|thumb|Shrub vegetation (with some [[cactus]]) in [[Webb County, Texas|Webb County]], Texas.]] [[File:Schlehenbusch.jpg|thumb|[[Prunus spinosa|Blackthorn shrub]] (''Prunus spinosa'') in the [[Vogelsberg Mountains|Vogelsberg]]]] [[File:Hortensie, blau.jpg|thumb|''[[Hydrangea macrophylla]]'']] [[File:Zaubernuss.jpg|thumb|Winter-flowering [[Witch-hazel]] (''Hamamelis'')]] [[File:Senecio angulatus by the coast.jpg|thumb|''[[Senecio angulatus]]'', a scrambling shrub by the sea (yellow-flowered).]] In [[botany]] and [[ecology]], a shrub is more specifically used to describe the particular [[Canopy (biology)|physical canopy structure]] or [[plant life-form]] of woody plants which are less than {{convert|8|m|ft}} high and usually multiple stems arising at or near the surface of the ground. For example, a descriptive system widely adopted in [[Australia]] is based on structural characteristics based on life-form, plus the height and amount of foliage cover of the tallest layer or [[Dominance (ecology)|dominant]] [[species]].<ref>Chris Paul (2024). [https://shrubradar.com/best-evergreen-shrubs-for-the-front-of-your-house The Best Evergreen Shrubs for Front Yards]. Shrubs Radar, a website all about shrubs.</ref> For shrubs that are {{convert|2|-|8|m|ft}} high, the following structural forms are categorized: * dense foliage cover (70β100%) β '''closed-shrubs''' * mid-dense foliage cover (30β70%) β '''open-shrubs''' * sparse foliage cover (10β30%) β '''tall shrubland''' * very sparse foliage cover (<10%) β '''tall open shrubland''' For shrubs less than {{convert|2|m|ft}} high, the following structural forms are categorized: * dense foliage cover (70β100%) β '''closed-[[heath (habitat)|heath]]''' or '''closed low shrubland'''β(''North America'') * mid-dense foliage cover (30β70%) β '''open-heath''' or '''mid-dense low shrubland'''β(''North America'') * sparse foliage cover (10β30%) β '''low shrubland''' * very sparse foliage cover (<10%) β '''low open shrubland''' == List == Those marked with * can also develop into tree form if in ideal conditions. {{div col begin|colwidth=20em}} ;A * ''[[Abelia]]'' (Abelia) * ''[[Acer (plant)|Acer]]'' (Maple) * * ''[[Actinidia]]'' (Actinidia) * ''[[Aloe]]'' (Aloe) * ''[[Aralia]]'' (Angelica Tree, [[Aralia spinosa|Hercules' Club]]) * * ''[[Arctostaphylos]]'' ([[Bearberry]], [[Manzanita]]) * * ''[[Aronia]]'' (Chokeberry) * ''[[Artemisia (genus)|Artemisia]]'' ([[Sagebrush]]) * ''[[Aucuba]]'' (Aucuba) ;B * ''[[Berberis]]'' (Barberry) * ''[[Bougainvillea]]'' (Bougainvillea) * ''[[Brugmansia]]'' (Angel's trumpet) * ''[[Buddleja]]'' (Butterfly bush) * ''[[Buxus]]'' (Box) * ;C * ''[[Calia]]'' (Mescalbean) * ''[[Callicarpa]]'' (Beautyberry) * * ''[[Callistemon]]'' (Bottlebrush) * * ''[[Calluna]]'' (Heather) * ''[[Calycanthus]]'' (Sweetshrub) * ''[[Camellia]]'' (Camellia, [[Tea]]) * * ''[[Caragana]]'' ([[Pea-tree]]) * * ''[[Carpenteria]]'' (Carpenteria) * ''[[Caryopteris]]'' ([[Blue Spiraea]]) * ''[[Cassiope]]'' ([[Moss-heather]]) * ''[[Ceanothus]]'' (Ceanothus) * * ''[[Celastrus]]'' (Staff vine) * * ''[[Ceratostigma]]'' ([[Hardy Plumbago]]) * ''[[Cercocarpus]]'' (Mountain-mahogany) * * ''[[Chaenomeles]]'' (Japanese Quince) * ''[[Chamaebatiaria]]'' (Fernbush) * ''[[Chamaedaphne]]'' (Leatherleaf) * ''[[Chimonanthus]]'' (Wintersweet) * ''[[Chionanthus]]'' (Fringe-tree) * * ''[[Choisya]]'' (Mexican-orange Blossom) * * ''[[Cistus]]'' (Rockrose) * ''[[Clerodendrum]]'' (Clerodendrum) * ''[[Clethra]]'' (Summersweet, Pepperbush) * * ''[[Clianthus]]'' ([[Glory Pea]]) * ''[[Colletia]]'' (Colletia) * ''[[Colutea]]'' (Bladder Senna) * ''[[Comptonia (plant)|Comptonia]]'' (Sweetfern) * ''[[Dogwood|Cornus]]'' (Dogwood) * * ''[[Corylopsis]]'' (Winter-hazel) * * ''[[Cotinus]]'' (Smoketree) * * ''[[Cotoneaster]]'' (Cotoneaster) * * ''[[Cowania (plant)|Cowania]]'' ([[Cliffrose]]) * ''[[Crataegus]]'' (Hawthorn) * * ''[[Crinodendron]]'' (Crinodendron) * * ''[[Cytisus]]'' and allied genera ([[Broom (shrub)|Broom]]) * ;D * ''[[Daboecia]]'' ([[Heath]]) * ''[[Danae (plant)|Danae]]'' (Alexandrian laurel) * ''[[Daphne (plant)|Daphne]]'' (Daphne) * ''[[Decaisnea]]'' (Decaisnea) * ''[[Dasiphora]]'' (Shrubby Cinquefoil) * ''[[Dendromecon]]'' (Tree poppy) * ''[[Desfontainea]]'' (Desfontainea) * ''[[Deutzia]]'' (Deutzia) * ''[[Diervilla]]'' (Bush honeysuckle) * ''[[Dipelta]]'' (Dipelta) * ''[[Dirca]]'' (Leatherwood) * ''[[Dracaena (plant)|Dracaena]]'' (Dragon tree) * * ''[[Drimys]]'' ([[Winter's Bark]]) * * ''[[Dryas (plant)|Dryas]]'' (Mountain Avens) ;E * ''[[Edgeworthia]]'' (Paper Bush) * * ''[[Elaeagnus]]'' ([[Elaeagnus]]) * * ''[[Embothrium]]'' ([[Chilean Firebush]]) * * ''[[Empetrum]]'' ([[Crowberry]]) * ''[[Enkianthus]]'' ([[Pagoda Bush]]) * ''[[Ephedra (genus)|Ephedra]]'' (Ephedra) * ''[[Epigaea]]'' ([[Trailing Arbutus]]) * ''[[Erica (plant)|Erica]]'' ([[Heath]]) * ''[[Eriobotrya]]'' (Loquat) * * ''[[Escallonia]]'' (Escallonia) * ''[[Eucryphia]]'' (Eucryphia) * * ''[[Euonymus]]'' (Spindle) * * ''[[Exochorda]]'' (Pearl Bush) ;F * ''[[Fabiana (plant)|Fabiana]]'' (Fabiana) * ''[[Fallugia]]'' (Apache Plume) * ''[[Fatsia]]'' (Fatsia) * ''[[Forsythia]]'' (Forsythia) * ''[[Fothergilla]]'' (Fothergilla) * ''[[Franklinia]]'' (Franklinia) * * ''[[Fremontodendron]]'' (Flannelbush) * ''[[Fuchsia]]'' (Fuchsia) * ;G * ''[[Garrya]]'' (Silk-tassel) * * ''[[Gaultheria]]'' ([[Salal]]) * ''[[Gaylussacia]]'' ([[Huckleberry]]) * ''[[Genista]]'' ([[Broom (shrub)|Broom]]) * * ''[[Gordonia (plant)|Gordonia]]'' ([[Loblolly-bay]]) * * ''[[Grevillea]]'' (Grevillea) * ''[[Griselinia]]'' (Griselinia) * ;H * ''[[Hakea]]'' (Hakea) * * ''[[Halesia]]'' ([[Silverbell]]) * * ''[[Halimium]]'' (Rockrose) * ''[[Hamamelis]]'' (Witch-hazel) * * ''[[Hebe (plant)|Hebe]]'' (Hebe) * ''[[Hedera]]'' (Ivy) * ''[[Helianthemum]]'' (Rockrose) * ''[[Hibiscus]]'' (Hibiscus) * * ''[[Hippophae]]'' (Sea-buckthorn) * * ''[[Hoheria]]'' (Lacebark) * * ''[[Holodiscus]]'' ([[Creambush]]) * ''[[Hudsonia]]'' (Hudsonia) * ''[[Hydrangea]]'' (Hydrangea) * ''[[Hypericum]]'' ([[Hypericum calycinum|Rose of Sharon]]) * ''[[Hyssopus (plant)|Hyssopus]]'' (Hyssop) ;I * ''[[Ilex]]'' ([[Holly]]) * * ''[[Illicium]]'' ([[Star Anise]]) * * ''[[Indigofera]]'' (Indigo) * ''[[Itea (plant)|Itea]]'' ([[Sweetspire]]) ;J * ''[[Jamesia]]'' (Cliffbush) * ''[[Jasminum]]'' (Jasmine) * ''[[Juniperus]]'' (Juniper) * ;K * ''[[Kalmia]]'' (Mountain-laurel) * ''[[Kerria (plant)|Kerria]]'' (Kerria) * ''[[Kolkwitzia]]'' ([[Beautybush|Beauty-bush]]) ;L * ''[[Lagerstroemia]]'' (Crape-myrtle) * * ''[[Lapageria]]'' (Copihue) * ''[[Lantana]]'' (Lantana) * ''[[Lavandula]]'' (Lavender) * ''[[Lavatera]]'' (Tree Mallow) * ''[[Ledum]]'' (Ledum) * ''[[Leitneria]]'' (Corkwood) * * ''[[Lespedeza]]'' (Bush Clover) * * ''[[Leptospermum]]'' (Manuka) * * ''[[Leucothoe (plant)|Leucothoe]]'' (Doghobble) * ''[[Leycesteria]]'' (Leycesteria) * ''[[Ligustrum]]'' (Privet) * * ''[[Lindera]]'' (Spicebush) * * ''[[Linnaea]]'' (Twinflower) * ''[[Lonicera]]'' (Honeysuckle) * ''[[Lupinus]]'' (Tree Lupin) * ''[[Lycium]]'' (Boxthorn) ;M * ''[[Magnolia]]'' (Magnolia) * ''[[Mahonia]]'' (Mahonia) * ''[[Malpighia]]'' ([[Acerola]]) * ''[[Menispermum]]'' (Moonseed) * ''[[Menziesia]]'' (Menziesia) * ''[[Mespilus]]'' ([[Medlar]]) * * ''[[Microcachrys]]'' (Microcachrys) * ''[[Myrica]]'' (Bayberry) * * ''[[Myricaria]]'' (Myricaria) * ''[[Myrtus]]'' and allied genera (Myrtle) * ;N * ''[[Neillia]]'' (Neillia) * ''[[Nerium]]'' (Oleander) ;O * ''[[Olearia]]'' (Daisy bush) * * ''[[Osmanthus]]'' (Osmanthus) ;P * ''[[Pachysandra]]'' (Pachysandra) * ''[[Peony|Paeonia]]'' (Tree-peony) * ''[[Persoonia]]'' (Geebungs) * ''[[Philadelphus]]'' (Mock orange) * * ''[[Phlomis]]'' ([[Jerusalem Sage]]) * ''[[Photinia]]'' (Photinia) * * ''[[Physocarpus]]'' (Ninebark) * * ''[[Pieris (plant)|Pieris]]'' (Pieris) * ''[[Pistacia]]'' ([[Pistachio]], [[Pistacia lentiscus|Mastic]]) * * ''[[Pittosporum]]'' (Pittosporum) * * ''[[Plumbago]]'' (Leadwort) * ''[[Polygala]]'' (Milkwort) * ''[[Poncirus]]'' * * ''[[Prunus]]'' (Cherry) * * ''[[Purshia]]'' ([[Antelope Bush]]) * ''[[Pyracantha]]'' (Firethorn) ;Q * ''[[Quassia]]'' (Quassia) * * ''[[Oak|Quercus]]'' (Oak) * * ''[[Quillaja]]'' ([[Quillay]]) * ''[[Quintinia]]'' ([[Tawheowheo]]) * ;R * ''[[Buckthorn|Rhamnus]]'' (Buckthorn) * * ''[[Rhododendron]]'' (Rhododendron, [[Azalea]]) * * ''[[Rhus]]'' (Sumac) * * ''[[Ribes]]'' (Currant, Gooseberry) * ''[[Romneya]]'' (Tree poppy) * ''[[Rose|Rosa]]'' (Rose) * ''[[Rosmarinus]]'' ([[Rosemary]]) * ''[[Rubus]]'' ([[Bramble]], [[Raspberry]], [[Salmonberry]], [[Rubus phoenicolasius|Wineberry]]) * ''[[Ruta]]'' ([[Rue]]) ;S * ''[[Sabia (plant)|Sabia]]'' * * ''[[Salix]]'' ([[Willow]]) * * ''[[Salvia]]'' (Sage) * ''[[Sambucus]]'' (Elder) * * ''[[Santolina]]'' ([[Lavender Cotton]]) * ''[[Sapindus]]'' (Soapberry) * * ''[[Senecio]]'' (Senecio) * ''[[Jojoba|Simmondsia]]'' (Jojoba) * ''[[Skimmia]]'' (Skimmia) * ''[[Smilax]]'' (Smilax) * ''[[Sophora]]'' ([[KΕwhai]]) * * ''[[Sorbaria]]'' (Sorbaria) * ''[[Spartium]]'' (Spanish Broom) * ''[[Spiraea]]'' (Spiraea) * * ''[[Staphylea]]'' (Bladdernut) * * ''[[Stephanandra]]'' (Stephanandra) * ''[[Styrax]]'' * * ''[[Symphoricarpos]]'' (Snowberry) * ''[[Syringa]]'' ([[Lilac]]) * ;T * ''[[Tamarix]]'' (Tamarix) * * ''[[Taxus]]'' (Yew) * * ''[[Waratah|Telopea]]'' (Waratah) * * ''[[Thuja]]'' cvs. (Arborvitae) * * ''[[Thymelaea]]'' * ''[[Thyme|Thymus]]'' (Thyme) * ''[[Trochodendron]]'' * ;U * ''[[Ulex]]'' (Gorse) * ''[[Ulmus pumila celer]]'' ([[Turkestan elm β Wonder Hedge]]) * ''[[Ungnadia]]'' (Mexican Buckeye) ;V * ''[[Vaccinium]]'' ([[Bilberry]], [[Blueberry]], [[Cranberry]]) * ''[[Verbesina centroboyacana]]'' * ''[[Vervain|Verbena]]'' (Vervain) * ''[[Viburnum]]'' (Viburnum) * * ''[[Vinca]]'' (Periwinkle) * ''[[Viscum]]'' ([[Mistletoe]]) ;W * ''[[Weigela]]'' (Weigela) ;X * ''[[Xanthoceras]]'' * ''[[Xanthorhiza]]'' (Yellowroot) * ''[[Xylosma]]'' ;Y * ''[[Yucca]]'' (Yucca, [[Yucca brevifolia|Joshua tree]]) * ;Z * ''[[Zanthoxylum]]'' * * ''[[Zauschneria]]'' * ''[[Zenobia (plant)|Zenobia]]'' * ''[[Ziziphus]]'' * {{div col end}} == References == {{Reflist}} {{Botany}} {{Authority control}} [[Category:Plants]] [[Category:Plant morphology]] [[Category:Shrubs| ]] [[Category:Lists of plants]] [[Category:Plant life-forms]] [[Category:Plants by habit]]
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