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Taproot
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{{Short description|Dominant root from which other roots sprout laterally}} {{About|the plant root system|other uses}} [[File:Root Systems.svg|thumb|The two types of root systems in plants. The fibrous-root system (A) is characterized by many roots with similar sizes. In contrast, plants that use the taproot system (B) grow a main root, with smaller roots branching off. The letters indicate where the root systems begin.]] A '''taproot''' is a large, central, and dominant [[root]] from which other roots sprout laterally. Typically a taproot is somewhat straight and very thick, is tapering in shape, and grows directly downward.<ref>{{cite web|url=http://www.hcs.ohio-state.edu/mg/manual/botany.htm|title=Botany Manual|publisher=[[Ohio State University]]|archive-url=https://web.archive.org/web/20040806065528/http://www.hcs.ohio-state.edu/mg/manual/botany.htm|archive-date=2004-08-06}}</ref> In some plants, such as the [[carrot]], the taproot is a [[storage organ]] so well developed that it has been cultivated as a vegetable. The '''taproot system''' contrasts with the adventitious- or [[fibrous-root system]] of plants with many branched roots, but many plants that grow a taproot during [[germination]] go on to develop branching root structures, although some that rely on the main root for storage may retain the dominant taproot for centuries—for example, ''[[Welwitschia]]''. == Description == [[File:Dandelion Blackwell 0136.jpg|thumb|A [[dandelion]] taproot (left) with the rest of the plant (right)]] [[Dicot]]s, one of the two divisions of [[flowering plant]]s (angiosperms), start with a taproot,<ref name="Mauseth2009">{{cite book|author=James D. Mauseth|author-link=James D. Mauseth|title=Botany: an introduction to plant biology|url=https://books.google.com/books?id=xPLGdYW9t5kC&pg=PA145|access-date=28 September 2010|year=2009|publisher=Jones & Bartlett Learning|isbn=978-0-7637-5345-0|pages=145–}}</ref> which is one main root forming from the enlarging [[radicle]] of the seed. The tap root can be persistent throughout the life of the plant but is most often replaced later in the plant's development by a fibrous root system.<ref name="Mauseth2009"/><ref name="BergBerg2007">{{cite book|author1=Linda Berg|author2=Linda R. Berg|title=Introductory Botany: Plants, People, and the Environment|url=https://books.google.com/books?id=I71WWH9ZmfsC&pg=PA112|access-date=28 September 2010|date=23 March 2007|publisher=Cengage Learning|isbn=978-0-534-46669-5|pages=112–}}</ref> A persistent taproot system forms when the radicle keeps growing and smaller lateral roots form along the taproot. The shape of taproots can vary but the typical shapes include: *Conical root: this type of root tuber is conical in shape, i.e. widest at the top and tapering steadily towards the bottom: e.g. [[carrot]]. *Fusiform root: this root is widest in the middle and tapers towards the top and the bottom: e.g. [[radish]]. *Napiform root: the root has a [[Spinning top|top]]-like appearance. It is very broad at the top and tapers suddenly like a tail at the bottom: e.g. [[turnip]]. [[File:Karotoj.jpg|thumb|The edible, orange part of the carrot is its taproot]] Many taproots are modified into [[storage organ]]s. Some plants with taproots: *[[Beetroot]] *[[Burdock]] *[[Carrot]] *[[Sugar beet]] *[[Dandelion]] *[[Parsley]] *[[Parsnip]] *[[Poppy mallow]] *[[Radish]] *[[Sagebrush]] *[[Turnip]] *[[Asclepias syriaca|Common milkweed]] * trees such as [[oak]]s, [[elm]]s, [[pine]]s and [[fir]]s == Development == Taproots develop from the [[radicle]] of a seed, forming the primary root. It branches off to secondary roots, which in turn branch to form tertiary roots. These may further branch to form rootlets. For most plants species the radicle dies some time after seed germination, causing the development of a fibrous root system, which lacks a main downward-growing root. Most [[tree]]s begin life with a taproot,<ref name="BergBerg2007"/> but after one to a few years the main root system changes to a wide-spreading fibrous root system with mainly horizontal-growing surface roots and only a few vertical, deep-anchoring roots. A typical mature tree 30–50 m tall has a root system that extends horizontally in all directions as far as the tree is tall or more, but as much as 100% of the roots are in the top 50 cm of soil. Soil characteristics strongly influence the architecture of taproots; for example, deep and rich soils favour the development of vertical taproots in many oak species such as ''[[Quercus kelloggii]]'', while clay soils promote the growth of multiple taproots.<ref>{{cite web |author=C. Michael Hogan |year=2008 |title=Quercus kelloggii |website=Globaltwitcher.com |editor=N. Stromberg |url=http://www.globaltwitcher.com/artspec_info.asp?thingid=82385 |access-date=2008-10-17 |url-status=dead |archive-url=https://web.archive.org/web/20090924051817/http://www.globaltwitcher.com/artspec_info.asp?thingid=82385 |archive-date=2009-09-24 }}</ref> ==Horticultural considerations== {{No footnotes|section|date=July 2020}} Many plants with taproots are difficult to [[transplanting|transplant]], or even to grow in containers, because the root tends to grow deep rapidly and in many species comparatively slight obstacles or damage to the taproot will stunt or kill the plant. Among weeds with taproots [[Taraxacum|dandelion]]s are typical; being deep-rooted, they are hard to uproot and if the taproot breaks off near the top, the part that stays in the ground often resprouts such that, for effective control, the taproot needs to be severed at least several centimetres below ground level. == Gallery == <gallery class="center" widths="200"> file:Buckeye6.jpg|A tree taproot file:Plant taproots.jpg|A taproot </gallery> == References == {{Reflist}} == External links == {{commons|Taproots}} *{{cite news |url=https://www.sciencedaily.com/releases/2006/01/060112035906.htm |date=2006-01-13 |work=Sciencedaily |title=Deep-rooted Plants Have Much Greater Impact On Climate Than Experts Thought |quote=The tap roots transfer [[rain]]water from the surface to reservoirs deep underground and redistribute water... increases [[photosynthesis]] and the evaporation of water... by 40 percent in the [[dry season]]... During the wet season, these plants can store as much as 10 percent of the annual [[Precipitation (meteorology)|precipitation]] as deep as 13 meters (43 feet) underground, to be tapped during the dry months... tree roots acting like pipes to allow water to shift around much faster than it could otherwise [[percolate]] through the [[soil]]. }} *[http://www.arboretum.fullerton.edu/grow/taproot.asp Fullerton Arboretum on taproots] {{Webarchive|url=https://web.archive.org/web/20080308052559/http://arboretum.fullerton.edu/grow/taproot.asp |date=2008-03-08 }} [[Category:Plant roots]]
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