Template:Short description {{#invoke:other uses|otheruses}} Template:About Template:Speciesbox The northern flicker or common flicker (Colaptes auratus) is a medium-sized bird of the woodpecker family. It is native to most of North America, parts of Central America, Cuba, and the Cayman Islands, and is one of the few woodpecker species that migrate. Over 100 common names for the northern flicker are known, including yellowhammer (not to be confused with the Eurasian yellowhammer (Emberiza citrinella)), clape, gaffer woodpecker, harry-wicket,<ref>{{#invoke:citation/CS1|citation |CitationClass=web }}</ref> heigh-ho, wake-up, walk-up, wick-up, yarrup, and gawker bird. Many of these names derive from attempts to imitate some of its calls. It is the state bird of Alabama (known by its colloquial name "yellowhammer").<ref>{{#invoke:citation/CS1|citation |CitationClass=web }}</ref>
TaxonomyEdit
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The English naturalist Mark Catesby described and illustrated the northern flicker in his book The Natural History of Carolina, Florida and the Bahama Islands which was published between 1729 and 1732. Catesby used the English name "Gold-winged Wood-pecker" and the Latin {{#invoke:Lang|lang}}.<ref>Template:Cite book</ref> When in 1758 the Swedish naturalist Carl Linnaeus updated his Systema Naturae for the tenth edition, he included the northern flicker, coined the binomial name Cuculus auratus and cited Catesby's book.<ref>Template:Cite book</ref>
The specific epithet {{#invoke:Lang|lang}} is a Latin word meaning "gilded" or "ornamented with gold".<ref>Template:Cite book</ref> The type locality is South Carolina.<ref>Template:Cite book</ref> The northern flicker is one of 13 extant New World woodpeckers now placed in the genus Colaptes that was introduced by the Irish zoologist Nicholas Aylward Vigors in 1825 with the northern flicker (Colaptes auratus) as the type species.<ref>Template:Cite journal</ref>
SubspeciesEdit
Ten subspecies are recognized, one of which is now extinct, though it may be invalid.<ref name="ITIS">{{#invoke:citation/CS1|citation |CitationClass=web }}</ref> The extant subspecies were at one time considered subspecies of two separate species called the yellow-shafted flicker (C. auratus, with four subspecies) and the red-shafted flicker (C. cafer, with six subspecies, five living and one extinct), but they commonly interbreed where their ranges overlap and are now considered one species by the American Ornithologists Union. This is an example of what is referred to in science as the species problem.
Yellow-shafted groupEdit
- The southern yellow-shafted flicker (C. a. auratus) resides in the southeastern United States from Florida to Virginia.<ref name="Pyle">Template:Cite book</ref> It is yellow under the tail and underwings and has yellow shafts on its primaries. It has a gray cap, a beige face, and a red bar at the nape of the neck. Males have a black mustache. Colaptes comes from the Greek verb colapt, meaning "to peck"; {{#invoke:Lang|lang}} is from the Latin root {{#invoke:Lang|lang}}, meaning "gold" or "golden", and refers to the bird's underwings. As the state bird of Alabama,<ref>{{#invoke:citation/CS1|citation
|CitationClass=web }}</ref> this subspecies is known by the common name "yellowhammer", a term that originated during the American Civil War to describe Confederate soldiers from Alabama.<ref>Template:Cite book</ref>
- The northern yellow-shafted flicker (C. a. luteus; formerly C. a. borealis) resides from central Alaska throughout most of Canada to southern Labrador, Newfoundland, and the northeastern United States.
- The Cuban yellow-shafted flicker (C. a. chrysocaulosus) is restricted to Cuba.
- The Grand Cayman yellow-shafted flicker (C. a. gundlachi) is restricted to Grand Cayman in the Cayman Islands. The subspecific epithet is named after Cuban naturalist Juan Gundlach.
Red-shafted groupEdit
- The western red-shafted flicker (C. a. cafer) resides in western North America. It is red under the tail and underwings and has red shafts on its primaries. It has a beige cap and a gray face. Males have a red mustache. The subspecific name cafer is the result of an error made in 1788 by the German systematist Johann Gmelin, who believed that its original habitat was in South Africa among the Xhosa people, then known as the "Kaffirs". As the origin of the subspecies designation is regarded as offensive by some, proposals to change the scientific name of this subspecies to C. a. lathami have been presented to the American Ornithological Society. The Society, in accordance with the rules governing scientific nomenclature, has as of September 2018 declined to support a change of the subspecific name, but may consult with the ICZN on the matter.<ref>{{#invoke:citation/CS1|citation
|CitationClass=web }}</ref><ref>Template:Cite journal</ref>
- The coastal red-shafted flicker (C. a. collaris) has a range that closely overlaps that of C. a. cafer, extending along much of the West Coast of North America from British Columbia to northwestern Mexico.
- The dwarf red-shafted flicker (C. a. nanus) resides in western Texas south to northeastern Mexico.
- The Mexican red-shafted flicker (C. a. mexicanus) resides in central and southern Mexico from Durango to San Luis Potosí and Oaxaca.
- The Guatemalan red-shafted flicker (C. a. mexicanoides) resides in the highlands of southern Mexico and Central America. It is considered by some authorities to be a separate species, the Guatemalan flicker (C. mexicanoides).
- The Guadalupe red-shafted flicker (C. a. rufipileus)† is extinct and was formerly restricted to Guadalupe Island, off the northwestern coast of Baja California, Mexico. It was last recorded in 1906. It may be invalid.<ref>{{#invoke:citation/CS1|citation
|CitationClass=web }}</ref> Individuals of an extant mainland red-shafted subspecies (which one is unknown) were observed breeding on Guadalupe Island in 1996.<ref>Template:Cite journal</ref>
- Southeastern Yellow-shafted Flicker, Kent County, DE, USA imported from iNaturalist photo 62634945.jpg
C. a. auratus, in Delaware
- Northern flicker pair.jpg
C. a. cafer female (left) and male (right), in Washington
- Northern flicker (Colaptes auratus chrysocaulosus) female.JPG
C. a. chrysocaulosus female, in Cuba
- Southwestern Red-shafted Flicker, Foothills Green, Fort Collins, CO 80526, USA imported from iNaturalist photo 452304844 (cropped).jpg
C. a. collaris male, in Colorado
- Grand Cayman Flicker, Cayman Islands, Cayman Islands, Cayman Islands, KY imported from iNaturalist photo 35488401 (cropped).jpg
C. a. gundlachi, in Grand Cayman
- Northern Yellow-shafted Flicker, Powderhorn Park, Minneapolis, MN 55407, USA imported from iNaturalist photo 437335546 (cropped).jpg
C. a. luteus male, in Minnesota
- Colaptes auratus mexicanoides 84221588.jpg
C. a. mexicanoides female, in Guatemala
- Colaptes rufipileus (Guadalupe flicker).jpg
†C. a. rufipileus male taxidermied specimen
DescriptionEdit
Adults are brown with black bars on the back and wings. A mid- to large-sized northern flicker measures Template:Convert in length and Template:Convert in wingspan.<ref name="CLO-AAB" /><ref>{{#invoke:citation/CS1|citation |CitationClass=web }}</ref> The body mass can vary from Template:Convert.<ref name="CRC">CRC Handbook of Avian Body Masses by John B. Dunning Jr. (Editor). CRC Press (1992), Template:ISBN.</ref> Among standard scientific measurements, the wing bone measures Template:Convert, the tail measures Template:Convert, the bill measures Template:Convert and the tarsus measures Template:Convert. The largest-bodied specimens are from the northern stretches of the species' range at the latitude of Alaska and Labrador, while the smallest specimens come from Grand Cayman Island.<ref>Template:Cite book</ref> A necklace-like black patch occupies the upper breast, while the lower breast and belly are beige with black spots. Males can be identified by a black (in the eastern part of the species' range) or red (in the western part) mustachial stripe at the base of the beak, while females lack this stripe. The tail is dark on top, transitioning to a white rump which is conspicuous in flight. Subspecific plumage is variable.
Call and flightEdit
Template:Multiple image This bird's call is a sustained laugh, ki ki ki ki, quite different from that of the pileated woodpecker (Dryocopus pileatus). One may also hear a constant knocking as they often drum on trees or even metal objects to declare territory. Like most woodpeckers, northern flickers drum on objects as a form of communication and territory defense. In such cases, the purpose is to make as loud a noise as possible, so woodpeckers sometimes drum on metal objects.
Like many woodpeckers, its flight is undulating. The repeated cycle of a quick succession of flaps followed by a pause creates an effect comparable to a roller coaster.
DietEdit
According to the Audubon field guide, "flickers are the only woodpeckers that frequently feed on the ground", probing with their beak, also sometimes catching insects in flight. Although they eat fruits, berries, seeds, and nuts, their primary food is insects. Ants alone can make up 45% of their diet. Other invertebrates eaten include flies, butterflies, moths, beetles, and snails. Northern flickers also eat berries and seeds, especially in winter, including those of poison ivy, poison oak, dogwood, sumac, wild cherry and grape, bayberries, hackberries, and elderberries, as well as sunflower and thistle seeds. Northern flickers often break into underground ant colonies to get at the nutritious larvae there, hammering at the soil the way other woodpeckers drill into wood. They have been observed breaking up cow dung to eat the insects living within. Their tongues can dart out Template:Convert beyond the end of the bill to catch prey.<ref name="CLO-AAB" /> The northern flicker is a natural predator of the European corn borer (Ostrinia nubilalis), an invasive species of moth that costs the U.S. agriculture industry more than $1 billion annually in crop losses and population control.<ref>{{#invoke:citation/CS1|citation |CitationClass=web }}</ref> As well as eating ants, northern flickers exhibit a behavior known as anting, in which they use the formic acid from the ants to assist in preening, as it is useful in keeping them free of parasites.
Influence of diet on offspringEdit
According to an article published in Ibis, the availability of food affects the coloration of feathers in northern flicker nestlings. The article focused on the correlation between melanin spots and carotenoid-based coloration on the wings of nestlings with food stress via indirect manipulation of brood size. The article found that there was a positive correlation between the quality of the nestlings' diet and T-cell-mediated immune response. T-cell-mediated immune response was found to be positively correlated with brightness of pigmentation in flight feathers, but not related to melanin spot intensity.<ref>{{#invoke:citation/CS1|citation |CitationClass=web }}</ref>
HabitatEdit
The northern flicker may be observed in open habitats near trees, including woodlands, edges, yards, and parks. In the western United States, one can find it in mountain forests all the way up to the tree line. The northern flicker generally nests in holes in trees like other woodpeckers. Occasionally, it has been found nesting in old, earthen burrows vacated by belted kingfishers (Megaceryle alcyon) or sand martins (Riparia riparia).<ref name="AAB-Overview">{{#invoke:citation/CS1|citation |CitationClass=web }}</ref> Both sexes help with the nest excavation. The entrance hole is about Template:Cvt in diameter, and the cavity is Template:Cvt deep. The cavity widens at the bottom to make room for the eggs and the incubating adult. Inside, the cavity is bare except for a bed of wood chips for the eggs and chicks to rest on.<ref name="Wild-Bird-Watching.com">{{#invoke:citation/CS1|citation |CitationClass=web }}</ref> Once the nestlings are about 17 days old, they begin clinging to the cavity wall rather than lying on the floor. They can create cavities inside homes, especially homes of stucco or weak wooden siding.<ref name="Wild-Bird-Watching.com" />
LifespanEdit
A study from 2006 examined the mortality rates of male and female northern flickers over a six-year period using capture-tag-recapture techniques. The researchers observed that only one to two birds out of every 300 adults were 7 or more years old. This observation data correlated well with a mortality model that predicted a 0.6% 7-year survival rate.<ref name="Fisher2006">Template:Cite journal</ref> The data also illustrated that there were no significant differences between male and female survival rates for the general population. The oldest yet known "yellow-shafted" northern flicker lived to be at least 9 years 2 months old, and the oldest yet known "red-shafted" northern flicker lived to be at least 8 years 9 months old.<ref name="CLO-AAB">{{#invoke:citation/CS1|citation |CitationClass=web }}</ref>
ReproductionEdit
The northern flicker's breeding habitat consists of forested areas across North America and as far south as Central America. It is a cavity nester that typically nests in trees, but may also use posts and birdhouses if sized and situated appropriately. It prefers to excavate its own home, although it may reuse and repair damaged or abandoned nests. Often the old nests are created by belted kingfishers or sand martins.<ref name="AAB-Overview"/> Abandoned northern flicker nests create habitats for other cavity nesters. The northern flicker is sometimes driven away from its nesting site by other cavity nesters like the common starling (Sturnus vulgaris).
The northern flicker commonly breeds during the months of February to July, depending upon the temperature of the area. During the breeding season, both mates will stay together. After the season, they do not stay together.<ref name="Wild-Bird-Watching.com"/> Before breeding season, one to two weeks are needed for a mated pair to build the nest. Male flickers find female flickers by head bobbing and their personal mating call.<ref name="Pappas">{{#invoke:citation/CS1|citation |CitationClass=web }}</ref> The common sounds the male bird makes towards a female is "woikawoikawoika", symbolizing their relationship to one another and other birds. If the call is used towards a male, it is a territorial sign.<ref name="Wild-Bird-Watching.com" /> The calls' pattern can be classified as flat and gradually rises into a loud noise. The call type is a chirp that drums and rattles.<ref name="www.audubon.org">{{#invoke:citation/CS1|citation |CitationClass=web }}</ref>
Additional territorial signs towards other males can be head swinging when in close proximity to another or repeatedly creating loud noises with their beaks. Commonly used objects are wood or metal for a louder sound.<ref name="AAB-Overview" /> While making the loud noises, they will spread their wings, moving them up and down, spreading their tail to flash their colorful underside.<ref name="www.audubon.org" /> The color of their feathers depends on the environment surrounding the bird.<ref>Template:Cite journal</ref> Currently, there is no direct correlation between the birds' colors and mate choice. Rather, it plays a bigger role in territory.<ref>{{#invoke:citation/CS1|citation |CitationClass=web }}</ref>
The northern flicker may also point its bill forward towards a competitor for territorial reasons.<ref name="Pappas" /> Juvenile northern flickers are often helpless against predators who enter the nest. The common predators are Cooper's hawks (Astur cooperii), sharp-shinned hawks (Accipter striatus), raccoons (Procyon lotor), squirrels, and snakes. Full grown northern flickers are preyed upon by larger birds and hunting birds.<ref name="Pappas" /> The entrance hole of their nest is roughly Template:Convert wide. The hole entrance is often facing east to southeast. On average, the northern flicker can have one to two clutches each breeding season.<ref name="Wild-Bird-Watching.com" /> A typical clutch consists of six to eight eggs whose shells are pure white with a smooth surface and high gloss. The eggs are the second-largest of the North American woodpecker species, exceeded only by the pileated woodpecker's. Incubation is by both sexes for about 11 to 12 days. Commonly the male will sit on the eggs overnight, and both the male and female will incubate the eggs during the day.<ref name="Wild-Bird-Watching.com" /> The young are fed by regurgitation and fledge about 25 to 28 days after hatching.
- Northern-flicker-males-territorial-display.jpg
Two males in a territorial display during spring
- Northern Flicker - 51159532100.jpg
In a dominance display, a northern flicker spreads its wings to show off its size and colors
- Flicker hole in CP (31848).jpg
Male guarding its nest cavity
- Northern-flicker-feeding.jpg
Adult feeding a juvenile at a nest cavity entrance
- Northern Flicker (m) carrying Fecal Sac from nest (53009673598).jpg
Male carrying a fecal sac away from the nest
Wintering and migrationEdit
Northern flickers are partial migrants, in which some southern populations are completely non-migratory.<ref name="Gow-2014">Template:Cite journal</ref> Those that do migrate tend to begin their spring migration towards the beginning of April and make their return between September and October.<ref>{{#invoke:citation/CS1|citation |CitationClass=web }}</ref> Individuals that breed farther north travel greater distances than their migratory southern conspecifics, often resulting in the convergence of northern and southern populations at wintering sites.<ref name="Gow-2014" /> This discrepancy likely arises from the northern flickers' ground foraging behavior, in which prey can only be found in snow-free locations.<ref name="Gow-2014" /> Furthermore, females tend to winter farther north than males, suggesting that parental investment and division of reproductive labour are key factors in determining individual migratory behavior.<ref name="Gow-2014" /> Rising temperatures resulting from anthropogenic climate change have been shown to trigger migration prematurely in Northern flickers, as well as many other migratory bird species as evidenced in Sherbrooke, Quebec.<ref>Template:Cite journal</ref>
Northern flickers are divided into eastern and western populations by the Rocky Mountains, with each population having a unique migratory pathway.<ref name="Gow-2014" /> Individuals breeding in the prairie provinces of Canada, the Dakotas, and surrounding U.S. states winter in Texas, Oklahoma, and Arkansas.<ref name="Gow-2014" /> Those breeding in southern Ontario and Michigan to New England winter from east Texas to the Carolinas, whereas those breeding in British Columbia and the Pacific Northwest winter from central California to Baja Peninsula, Mexico.<ref name="Gow-2014" />
During migration, northern flickers may form flocks.<ref name="Gow-2014-2">Template:Cite journal</ref> Additionally, the species' propensity for roosting in cavities is not mitigated during migration. On average, 75% of individuals spend their nights in a cavity during migration, even in completely unknown locations.<ref name="Gow-2014-2" /> Northern flickers demonstrate a high rate of nest cavity re-use, as opposed to excavating new cavities each year.<ref>Template:Cite journal</ref> Furthermore, breeding individuals exhibit intense site fidelity, with pairs consistently returning to the specific nest cavity they used in the previous year.<ref>Template:Cite journal</ref>
GalleryEdit
- Northern Flicker on Seedskadee National Wildlife Refuge (25906541634).jpg
A northern flicker at a tree in the Seedskadee National Wildlife Refuge
- Southeastern Yellow-shafted Flicker, Overton Park, TN, US imported from iNaturalist photo 3008122.jpg
C. a. auratus
- 'Colaptes mexicanus' by Edward Hargitt, 1889.jpg
Painting of "Colaptes mexicanus" by Edward Hargitt, 1889
- Northern Yellow-shafted Flicker, Powderhorn Park, Minneapolis, MN 55407, USA imported from iNaturalist photo 437335514.jpg
C. a. luteus, in Minnesota
- Northern Yellow-shafted Flicker, Powderhorn Park, Minneapolis, MN 55407, USA imported from iNaturalist photo 437335534.jpg
C. a. luteus, in Minnesota
- Grand Cayman Flicker, Cayman Islands, Cayman Islands, Cayman Islands, KY imported from iNaturalist photo 35488379.jpg
C. a. gundlachi, in Grand Cayman
- Southwestern Red-shafted Flicker, Bernardo Waterfowl Area, Bosque, NM imported from iNaturalist photo 348461362 (cropped).jpg
C. a. collaris, in New Mexico
- Northern Yellow-shafted Flicker, Yolo County, CA, USA imported from iNaturalist photo 341176313 (cropped).jpg
C. a. luteus, in California
ReferencesEdit
External linksEdit
Template:Sister project Template:Sister project
- Northern Flicker - Colaptes auratus - USGS Patuxent Bird Identification InfoCenter
- A sample of the call of a Northern Flicker by the USGS (mp3)
- Template:Usurped (for Antigua, Cayman Islands, Cuba, Saint Pierre and Miquelon, United States) (imperfect Range Map) at bird-stamps.org
- Template:EBirdSpecies
- Template:VIREO
- Calls of the northern flicker at Animal Diversity Web
- Northern (Yellow-shafted) Flicker Bird Sound at Florida Museum of Natural History