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The biosphere (Template:Etymology), also called the ecosphere (Template:Etymology), is the worldwide sum of all ecosystems. It can also be termed the zone of life on the Earth. The biosphere (which is technically a spherical shell) is virtually a closed system with regard to matter,<ref name="ColumbiaEncyc">"Biosphere" in The Columbia Encyclopedia, 6th ed. (2004) Columbia University Press.</ref> with minimal inputs and outputs. Regarding energy, it is an open system, with photosynthesis capturing solar energy at a rate of around 100 terawatts.<ref>Template:Cite journal</ref> By the most general biophysiological definition, the biosphere is the global ecological system integrating all living beings and their relationships, including their interaction with the elements of the lithosphere, cryosphere, hydrosphere, and atmosphere. The biosphere is postulated to have evolved, beginning with a process of biopoiesis (life created naturally from Template:Nowrap matter, such as simple organic compounds) or biogenesis (life created from living matter), at least some 3.5 billion years ago.<ref name="Campbell 2006">Template:Cite book</ref><ref name="NYT-20131003">Template:Cite news</ref>
In a general sense, biospheres are any closed, self-regulating systems containing ecosystems. This includes artificial biospheres such as Template:Nowrap and Template:Nowrap, and potentially ones on other planets or moons.<ref name="webdictionary.co.uk">{{#invoke:citation/CS1|citation |CitationClass=web }}</ref>
Origin and use of the termEdit
The term "biosphere" was coined in 1875 by geologist Eduard Suess, who defined it as the place on Earth's surface where life dwells.<ref>Suess, E. (1875) Die Entstehung Der Alpen [The Origin of the Alps]. Vienna: W. Braunmuller.</ref>
While the concept has a geological origin, it is an indication of the effect of both Charles Darwin and Matthew F. Maury on the Earth sciences. The biosphere's ecological context comes from the 1920s (see Vladimir I. Vernadsky), preceding the 1935 introduction of the term "ecosystem" by Sir Arthur Tansley (see ecology history). Vernadsky defined ecology as the science of the biosphere. It is an interdisciplinary concept for integrating astronomy, geophysics, meteorology, biogeography, evolution, geology, geochemistry, hydrology and, generally speaking, all life and Earth sciences.
Narrow definitionEdit
Geochemists define the biosphere as being the total sum of living organisms (the "biomass" or "biota" as referred to by biologists and ecologists). In this sense, the biosphere is but one of four separate components of the geochemical model, the other three being geosphere, hydrosphere, and atmosphere. When these four component spheres are combined into one system, it is known as the ecosphere. This term was coined during the 1960s and encompasses both biological and physical components of the planet.<ref>Template:Cite book</ref>
The Second International Conference on Closed Life Systems defined biospherics as the science and technology of analogs and models of Earth's biosphere; i.e., artificial Earth-like biospheres.<ref>Template:Cite book</ref> Others may include the creation of artificial non-Earth biospheres—for example, human-centered biospheres or a native Martian biosphere—as part of the topic of biospherics.<ref>Template:Cite journal</ref>
Earth's biosphereEdit
OverviewEdit
Currently, the total number of living cells on the Earth is estimated to be 1030; the total number since the beginning of Earth, as 1040, and the total number for the entire time of a habitable planet Earth as 1041.<ref name="NYT-20231201">Template:Cite news</ref><ref name="CB-20231106">Template:Cite journal</ref> This is much larger than the total number of estimated stars (and Earth-like planets) in the observable universe as 1024, a number which is more than all the grains of beach sand on planet Earth;<ref name="ESA-2020">{{#invoke:citation/CS1|citation |CitationClass=web }}</ref><ref name="SWIN-20020201">{{#invoke:citation/CS1|citation |CitationClass=web }}</ref><ref name="CNET-20150319">Template:Cite news</ref><ref name="MNRAS-20150313">Template:Cite journal</ref> but less than the total number of atoms estimated in the observable universe as 1082;<ref name="LS-20210711">Template:Cite news</ref> and the estimated total number of stars in an inflationary universe (observed and unobserved), as 10100.<ref name="SR-20200203">Template:Cite journal</ref>
AgeEdit
The earliest evidence for life on Earth includes biogenic graphite found in 3.7 billion-year-old metasedimentary rocks from Western Greenland<ref name="NG-20131208">Template:Cite journal</ref> and microbial mat fossils found in 3.48 billion-year-old sandstone from Western Australia.<ref name="AP-20131113">Template:Cite news</ref><ref name="AST-20131108">Template:Cite journal</ref> More recently, in 2015, "remains of biotic life" were found in 4.1 billion-year-old rocks in Western Australia.<ref name="AP-20151019">Template:Cite news</ref><ref name="PNAS-20151014-pdf">Template:Cite journal Early edition, published online before print.</ref> In 2017, putative fossilized microorganisms (or microfossils) were announced to have been discovered in hydrothermal vent precipitates in the Nuvvuagittuq Belt of Quebec, Canada that were as old as 4.28 billion years, the oldest record of life on earth, suggesting "an almost instantaneous emergence of life" after ocean formation 4.4 billion years ago, and not long after the formation of the Earth 4.54 billion years ago.<ref name="NAT-20170301">Template:Cite journal</ref><ref name="NYT-20170301">Template:Cite news</ref><ref name="BBC-20170301">{{#invoke:citation/CS1|citation |CitationClass=web }}</ref><ref name="4.3b oldest">Template:Cite news</ref> According to biologist Stephen Blair Hedges, "If life arose relatively quickly on Earth ... then it could be common in the universe."<ref name="AP-20151019" />
ExtentEdit
Every part of the planet, from the polar ice caps to the equator, features life of some kind. Recent advances in microbiology have demonstrated that microbes live deep beneath the Earth's terrestrial surface and that the total mass of microbial life in so-called "uninhabitable zones" may, in biomass, exceed all animal and plant life on the surface. The actual thickness of the biosphere on Earth is difficult to measure. Birds typically fly at altitudes as high as Template:Convert and fish live as much as Template:Convert underwater in the Puerto Rico Trench.<ref name="Campbell 2006" />
There are more extreme examples for life on the planet: Rüppell's vulture has been found at altitudes of Template:Convert; bar-headed geese migrate at altitudes of at least Template:Convert; yaks live at elevations as high as Template:Convert above sea level; mountain goats live up to Template:Convert. Herbivorous animals at these elevations depend on lichens, grasses, and herbs.
Life forms live in every part of the Earth's biosphere, including soil, hot springs, inside rocks at least Template:Convert deep underground, and at least Template:Convert high in the atmosphere.<ref name="SD-19980625-UG">{{#invoke:citation/CS1|citation |CitationClass=web }}</ref><ref name="ABM-20150112">{{#invoke:citation/CS1|citation |CitationClass=web }}</ref><ref name="BBC-20151124">{{#invoke:citation/CS1|citation |CitationClass=web }}</ref> Marine life under many forms has been found in the deepest reaches of the world ocean while much of the deep sea remains to be explored.<ref>Template:Cite journal[1]</ref>
Under certain test conditions, microorganisms have been observed to survive the vacuum of outer space.<ref name=Dose>Template:Cite journal</ref><ref name='Horneck'>Template:Cite journal</ref> The total amount of soil and subsurface bacterial carbon is estimated as 5 × 1017 g.<ref name="SD-19980625-UG" /> The mass of prokaryote microorganisms—which includes bacteria and archaea, but not the nucleated eukaryote microorganisms—may be as much as 0.8 trillion tons of carbon (of the total biosphere mass, estimated at between 1 and 4 trillion tons).<ref name="AGCI-2014">{{#invoke:citation/CS1|citation |CitationClass=web }}</ref> Barophilic marine microbes have been found at more than a depth of Template:Convert in the Mariana Trench, the deepest spot in the Earth's oceans.<ref>Template:Cite journal</ref> In fact, single-celled life forms have been found in the deepest part of the Mariana Trench, by the Challenger Deep, at depths of Template:Convert.<ref>{{#invoke:citation/CS1|citation |CitationClass=web }}</ref><ref name="LS-20130317">{{#invoke:citation/CS1|citation |CitationClass=web }}</ref><ref name="NG-20130317">Template:Cite journal</ref> Other researchers reported related studies that microorganisms thrive inside rocks up to Template:Convert below the sea floor under Template:Convert of ocean off the coast of the northwestern United States,<ref name="LS-20130317" /><ref name="LS-20130314">{{#invoke:citation/CS1|citation |CitationClass=web }}</ref> as well as Template:Convert beneath the seabed off Japan.<ref name="BBC-20141215-RM">Template:Cite news</ref> Culturable thermophilic microbes have been extracted from cores drilled more than Template:Convert into the Earth's crust in Sweden,<ref name="Szewzyketal1994">Template:Cite journal</ref> from rocks between Template:Convert. Temperature increases with increasing depth into the Earth's crust. The rate at which the temperature increases depends on many factors, including the type of crust (continental vs. oceanic), rock type, geographic location, etc. The greatest known temperature at which microbial life can exist is Template:Convert (Methanopyrus kandleri Strain 116). It is likely that the limit of life in the "deep biosphere" is defined by temperature rather than absolute depth.Template:Citation needed On 20 August 2014, scientists confirmed the existence of microorganisms living Template:Convert below the ice of Antarctica.<ref name="NAT-20140820">Template:Cite journal</ref><ref name="FRB-20140820">{{#invoke:citation/CS1|citation |CitationClass=web }}</ref>
Earth's biosphere is divided into several biomes, inhabited by fairly similar flora and fauna. On land, biomes are separated primarily by latitude. Terrestrial biomes lying within the Arctic and Antarctic Circles are relatively barren of plant and animal life. In contrast, most of the more populous biomes lie near the equator.
Annual variationEdit
Artificial biospheresEdit
Experimental biospheres, also called closed ecological systems, have been created to study ecosystems and the potential for supporting life outside the Earth. These include spacecraft and the following terrestrial laboratories:
- Biosphere 2 in Arizona, United States, 3.15 acres (13,000 m2).
- BIOS-1, BIOS-2 and BIOS-3 at the Institute of Biophysics in Krasnoyarsk, Siberia, in what was then the Soviet Union.<ref>Template:Cite journal</ref>
- Biosphere J (CEEF, Closed Ecology Experiment Facilities), an experiment in Japan.<ref>Template:Cite journal</ref><ref>{{#invoke:citation/CS1|citation
|CitationClass=web }}</ref>
- Micro-Ecological Life Support System Alternative (MELiSSA) at Autonomous University of Barcelona
Extraterrestrial biospheresEdit
No biospheres have been detected beyond the Earth; therefore, the existence of extraterrestrial biospheres remains hypothetical. The rare Earth hypothesis suggests they should be very rare, save ones composed of microbial life only.<ref>Template:Cite book</ref> On the other hand, Earth analogs may be quite numerous, at least in the Milky Way galaxy, given the large number of planets.<ref>Template:Cite news</ref> Three of the planets discovered orbiting TRAPPIST-1 could possibly contain biospheres.<ref>Template:Cite news</ref> Given limited understanding of abiogenesis, it is currently unknown what percentage of these planets actually develop biospheres.
Based on observations by the Kepler Space Telescope team, it has been calculated that provided the probability of abiogenesis is higher than 1 to 1000, the closest alien biosphere should be within 100 light-years from the Earth.<ref>Amri Wandel, On the abundance of extraterrestrial life after the Kepler mission Template:Webarchive</ref>
It is also possible that artificial biospheres will be created in the future, for example with the terraforming of Mars.<ref>Template:Cite book</ref>
See alsoEdit
Template:Portal Template:Div col
- Biosphere model
- Climate system
- Cryosphere
- Habitable zone
- Homeostasis
- Life-support system
- Man and the Biosphere Programme
- Montreal Biosphere
- Noosphere
- Rare biosphere
- Shadow biosphere
- Soil biomantle
- Thomas Gold
- Wardian case
- Winogradsky column
ReferencesEdit
Further readingEdit
- The Biosphere (A Scientific American Book), San Francisco, W.H. Freeman and Co., 1970, Template:ISBN. This book, originally the December 1970 Scientific American issue, covers virtually every major concern and concept since debated regarding materials and energy resources (including solar energy), population trends, and environmental degradation (including global warming).
External linksEdit
- Article on the Biosphere at Encyclopedia of Earth
- GLOBIO.info, an ongoing programme to map the past, current and future impacts of human activities on the biosphere
- Paul Crutzen Interview, freeview video of Paul Crutzen Nobel Laureate for his work on decomposition of ozone talking to Harry Kroto Nobel Laureate by the Vega Science Trust.
- Atlas of the Biosphere
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