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Typha Template:IPAc-en is a genus of about 30 species of monocotyledonous flowering plants in the family Typhaceae. These plants have a variety of common names, in British English as bulrush<ref name = "Streeter">Streeter D, Hart-Davies C, Hardcastle A, Cole F, Harper L. 2009. Collins Flower Guide. Harper Collins Template:ISBN</ref> or (mainly historically) reedmace,<ref>Clegg, J. (1986). Observer's Book of Pond Life. Frederick Warne, London. 460 p.</ref> in American English as cattail,<ref>Template:PLANTS</ref> or punks, in Australia as cumbungi or bulrush, in Canada as bulrush or cattail, and in New Zealand as raupō, bullrush,<ref>{{#invoke:citation/CS1|citation |CitationClass=web }}</ref> cattail or reed. Other taxa of plants may be known as bulrush, including some sedges in Scirpus and related genera.
The genus is largely distributed in the Northern Hemisphere, where it is found in a variety of wetland habitats.
The rhizomes are edible, though at least some species are known to accumulate toxins and so must first undergo treatment before being eaten.<ref name="State of Victoria">{{#invoke:citation/CS1|citation |CitationClass=web }}</ref> Evidence of preserved starch grains on grinding stones suggests they were already eaten in Europe 30,000 years ago.<ref name="Revedin">Template:Cite journal</ref>
DescriptionEdit
Typha are aquatic or semi-aquatic, rhizomatous, herbaceous perennial plants.<ref name=Stace>Template:Cite book</ref>Template:Rp The leaves are glabrous (hairless), linear, alternate and mostly basal on a simple, jointless stem that bears the flowering spikes. The plants are monoecious, with unisexual flowers that develop in dense racemes. The numerous male flowers form a narrow spike at the top of the vertical stem. Each male (staminate) flower is reduced to a pair of stamens and hairs, and withers once the pollen is shed. Large numbers of tiny female flowers form a dense, sausage-shaped spike on the stem below the male spike. In larger species this can be up to Template:Convert long and Template:Convert thick. The seeds are minute, Template:Convert long, and attached to fine hairs. When ripe, the heads disintegrate into a cottony fluff from which the seeds disperse by wind.
Fruits of Typha have been found as long ago as 69 MYA in modern Central Europe.<ref>Template:Cite journal</ref>
General ecologyEdit
Typha are often among the first wetland plants to colonize areas of newly exposed wet mud, with their abundant wind-dispersed seeds. Buried seeds can survive in the soil for long periods of time.<ref>van der Valk, A. G., and Davis, C. B. (1976). The seed banks of prairie glacial marshes. Canadian Journal of Botany 54, 1832–8.</ref> They germinate best with sunlight and fluctuating temperatures, which is typical of many wetland plants that regenerate on mud flats.<ref>Shipley, B., et al. (1989). Regeneration and establishment strategies of emergent macrophytes. Journal of Ecology 77, 1093–1110.</ref> The plants also spread by rhizomes, forming large, interconnected stands.
Typha are considered to be dominant competitors in wetlands in many areas, and they often exclude other plants with their dense canopy.<ref name=Keddy>Template:Cite book</ref> In the bays of the Great Lakes, for example, they are among the most abundant wetland plants. Different species of cattails are adapted to different water depths.<ref>Grace, J. B. and Wetzel, R. G. (1981). Habitat partitioning and competitive displacement in cattails (Typha): experimental field studies. The American Naturalist 118: 463–74.</ref>
Well-developed aerenchyma make the plants tolerant of submersion. Even the dead stalks are capable of transmitting oxygen to the rooting zone.
Although Typha are native wetland plants, they can be aggressive in their competition with other native species.<ref>Oudhia, P. (1999). Allelopathic TEMPeffects of Typha angustata on germination and seedling vigour of winter maize and rice. Agric. Sci. Digest 19(4): 285-286.</ref> They have been problematic in many regions in North America, from the Great Lakes to the Everglades.<ref name=Keddy/> Native sedges are displaced and wet meadows shrink, likely as a response to altered hydrology of the wetlands and increased nutrient levels. An introduced or hybrid species may be contributing to the problem.<ref>Boers, A. M., et al. (2007). Typha × glauca dominance and extended hydroperiod constrain restoration of wetland diversity. Ecological Engineering 29, 232–44.</ref> Control is difficult. The most successful strategy appears to be mowing or burning to remove the aerenchymous stalks, followed by prolonged flooding.<ref>Kaminski, R. M., et al. (1985). Control of cattail and bulrush by cutting and flooding. In: Coastal Wetlands, eds. H. H. Prince and F. M. D’Itri, pp. 253–62. Chelsea, MI: Lewis Publishers.</ref> It may be more important to prevent invasion by preserving water level fluctuations, including periods of drought, and to maintain infertile conditions.<ref name=Keddy/>
Typha are frequently eaten by wetland mammals such as muskrats, which also use them to construct feeding platforms and dens, thereby also providing nesting and resting places for waterfowl.<ref>Global Invasive Species Database: "Uses"- Retrieved 2017-03-20</ref>
Accepted species and natural hybridsEdit
The following species and hybrids are currently accepted:<ref>{{#invoke:citation/CS1|citation |CitationClass=web }}</ref> Template:Columns-list
The most widespread species is Typha latifolia, which is distributed across the entire temperate northern hemisphere. It has also been introduced to Australia. T. angustifolia is nearly as widespread, but does not extend as far north; it may be introduced and invasive in North America. T. domingensis has a more southern American distribution, and it occurs in Australia. T. orientalis is widespread in Asia, Australia, and New Zealand. T. laxmannii, T. minima, and T. shuttleworthii are largely restricted to Asia and southern Europe.
UsesEdit
CulinaryEdit
Many parts of the Typha plant are edible to humans. Before the plant flowers, the tender inside of the shoots can be squeezed out and eaten raw or cooked.<ref>Template:Cite book</ref> The starchy rhizomes are nutritious with a protein content comparable to that of maize or rice.<ref name=Morton>Template:Cite journal</ref> They can be processed into a flour with 266 kcal per 100 grams,<ref name="Revedin"/> and are most often harvested from late autumn to early spring. They are fibrous, and the starch must be scraped or sucked from the tough fibers.<ref name=nova/> Baby shoots emerging from the rhizomes, which are sometimes subterranean, can be picked and eaten raw. Also underground is a carbohydrate lump which can be peeled and eaten raw or cooked like a potato.<ref name=Nyerges40>Template:Cite book</ref> The plant is one championed by survival experts because various parts can be eaten throughout the year. Plants growing in polluted water can accumulate lead and pesticide residues in their rhizomes, and these should not be eaten.<ref name=nova>Template:Cite book</ref>
The rind of young stems can be peeled off, and the tender white heart inside can be eaten raw or boiled and eaten like asparagus.<ref>Template:Cite book</ref> This food has been popular among the Cossacks in Ukraine, and has been called "Cossack asparagus".<ref>Template:Cite journal</ref> The leaf bases can be eaten raw or cooked, especially in late spring when they are young and tender. In early summer the sheath can be removed from the developing green flower spike, which can then be boiled and eaten like corn on the cob.<ref>Template:Cite book</ref> In mid-summer when the male flowers are mature, the pollen can be collected and used as a flour supplement or thickener; the Māori of New Zealand have a special bread called pungapunga made from the pollen of T. orientalis.<ref>Template:Cite journal</ref><ref>Raupo or Bulrush (Typha orientalis). Tai Awatea. Accessed 15 December 2011.</ref>
AgricultureEdit
The seeds have a high linoleic acid content and can be used to feed cattle and chickens.<ref>Template:Cite journal</ref> They can also be found in African countries like Ghana.
Harvesting cattail removes nutrients from the wetland that would otherwise return via the decomposition of decaying plant matter.<ref>Template:Cite journal</ref> Floating mats of cattails remove nutrients from eutrophied bodies of freshwater.<ref>{{#invoke:citation/CS1|citation |CitationClass=web }}</ref>
Building materialEdit
For local native tribes around Lake Titicaca in Peru and Bolivia, Typha were among the most important plants and every part of the plant had multiple uses. For example, they were used to construct rafts and other boats.<ref name=Morton/>
During World War II, the United States Navy used the down of Typha as a substitute for kapok in life vests and aviation jackets. Tests showed that even after 100 hours of submersion, the buoyancy was still effective.<ref>Template:Cite book</ref>
Typha are used as thermal insulation in buildings<ref>Template:Cite journal</ref> as an organic alternative to conventional insulating materials such as glass wool or stone wool.
PaperEdit
Typha stems and leaves can be used to make paper. It is strong with a heavy texture and it is hard to bleach, so it is not suitable for industrial production of graphical paper. In 1853, considerable amounts of cattail paper were produced in New York, due to a shortage of raw materials.<ref>Template:Cite book</ref> In 1948, French scientists tested methods for annual harvesting of the leaves. Because of the high cost, these methods were abandoned and no further research was done.<ref name=Morton/> Today Typha is used to make decorative paper.<ref>Template:Cite journal</ref><ref>Template:Cite journal</ref>
FiberEdit
Fibers up to 4 meters long can be obtained from the stems when they are treated mechanically or chemically with sodium hydroxide. The stem fibers resemble jute and can be used to produce raw textiles. The leaf fibers can be used as an alternative to cotton and linen in clothing. The yield of leaf fiber is 30 to 40 percent and Typha glauca can produce 7 to 10 tons per hectare annually.<ref name=Morton/>
BiofuelEdit
Typha can be used as a source of starch to produce ethanol. Because of their high productivity in northern latitudes, Typha are considered to be a bioenergy crop.<ref>Template:Cite journal</ref>
OtherEdit
The seed hairs were used by some indigenous peoples of the AmericasTemplate:Which as tinder for starting fires. Some tribes also used Typha down to line moccasins, and for bedding, diapers, baby powder, and cradleboards. One Native American word for Typha meant "fruit for papoose's bed".Template:Citation needed Typha down is still used in some areas to stuff clothing items and pillows. Typha can be dipped in wax or fat and then lit as a candle, the stem serving as a wick. Without the use of wax or fat it will smolder slowly, somewhat like incense, and may repel insects. Template:Citation needed
The flower stalks can be made into chopsticks. The leaves can be treated to weave into baskets, mats, or sandals.<ref name=Nyerges40/> The rushes are harvested and the leaves often dried for later use in chair seats. Re-wetted, the leaves are twisted and wrapped around the chair rungs to form a densely woven seat that is then stuffed (usually with the left over rush).
Small-scale experiments have indicated that Typha are able to remove arsenic from drinking water.<ref>Template:Cite journal</ref><ref>Template:Cite journal</ref> The boiled rootstocks have been used as a diuretic for increasing urination, or mashed to make a jelly-like paste for sores, boils, wounds, burns, scabs, and smallpox pustules.<ref>Template:Cite book</ref>
Cattail pollen is used as a banker source of food for predatory insects and mites (such as Amblyseius swirskii) in greenhouses.<ref>{{#invoke:citation/CS1|citation |CitationClass=web }}</ref>
The cattail, or, as it is commonly referred to in the American Midwest, the sausage tail, has been the subject of multiple artist renditions, gaining popularity in the mid-twentieth century. The term, sausage tail, derives from the similarity that cattails have with sausages, a name given to the plant by the Midwest Polish community, which had noticed a striking similarity between the plant and a common Polish dish, kiełbasa.Template:Cn
ReferencesEdit
External linksEdit
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