Carpinus betulus
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Carpinus betulus, the European or common hornbeam, is a species of tree in the birch family Betulaceae, native to Western Asia and central, eastern, and southern Europe, including southern England.<ref name=nhm/> It requires a warm climate for good growth, and occurs only at elevations up to Template:Convert. It grows in mixed stands with oak, and in some areas beech, and is also a common tree in scree forests. Hornbeam was also known as yoke elm.<ref name="Perthensis">Template:Cite book</ref> Together with Carpinus orientalis and Carpinus austrobalcanica, it is one of the three hornbeams found in Europe.
DescriptionEdit
It is a deciduous small to medium-size tree reaching heights of Template:Convert, rarely Template:Convert, and often has a fluted and crooked trunk. The bark is smooth and greenish-grey, even in old trees. The buds, unlike those of the beech, are Template:Convert long at the most, and pressed close to the twig. The leaves are alternate, Template:Convert long, with prominent veins giving a distinctive corrugated texture, and a serrated margin. It is monoecious, and the wind-pollinated male and female catkins appear in early summer after the leaves. The fruit is a small Template:Convert long nut, partially surrounded by a three-pointed leafy involucre Template:Convert long; it matures in autumn.Template:Citation needed
Distribution
Hornbeam is considered native from Western Asia and throughout Europe.<ref name=nhm>{{#invoke:citation/CS1|citation |CitationClass=web }}</ref> The species prefers a warm climate, and only naturally occurs below Template:Convert in elevation. It is a common tree in scree forests.<ref name="Perthensis"/>
Hornbeam was frequently coppiced and pollarded in the past in England. It is still infrequently managed using these traditional methods, but mainly for non-commercial conservation purposes. As a woodland tree traditionally managed in this way, it is particularly frequent in the ancient woodlands of south Essex, Hertfordshire and north Kent where it typically occupies more than half of most ancient woods and wood pastures.<ref name=Rackham>Template:Cite book</ref>
There are a number of notable forests where C. betulus is a dominant tree species, among which include Epping Forest in the United Kingdom, and Halltorps hage in Öland, Sweden.
Fossil recordEdit
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Three fossil fruits of Carpinus betulus have been extracted from borehole samples of the Middle Miocene fresh water deposits in Nowy Sacz Basin, West Carpathians, Poland.<ref>Łańcucka-Środoniowa M.: Macroscopic plant remains from the freshwater Miocene of the Nowy Sącz Basin (West Carpathians, Poland) [Szczątki makroskopowe roślin z miocenu słodkowodnego Kotliny Sądeckiej (Karpaty Zachodnie, Polska)]. Acta Palaeobotanica 1979 20 (1): 3-117.</ref>
EcologyEdit
In England, trees appear to prefer soils with a pH from 3.6 to 4.6 but tolerate up to 7.6. They are found on soils with moderate clay content and avoid soils with particularly high or low clay content.<ref name=Rackham/> Carpinus betulus likes full sun or partial shade,<ref name=rhs/> moderate soil fertility and moisture. It has a shallow, wide-spreading root system and is marked by the production of stump sprouts when cut back.Template:Citation needed
The seeds often do not germinate until the spring of the second year after sowing. The hornbeam is a prolific seeder and is marked by vigorous natural regeneration.Template:Citation needed
Associated speciesEdit
Hornbeam grows in mixed stands with oak, and in some areas beech.<ref name="Perthensis"/> The leaves provide food for some animals, including Lepidoptera such as the case-bearer moth Coleophora anatipennella.Template:Citation needed
This tree has been associated with the poisonous mushroom Amanita phalloides, better known as the death-cap mushroom, which grow around the trunk after hornbeams mature. When ingested, death-cap mushrooms can cause extreme medical conditions and death. Death-cap mushrooms can be mistaken for other edible mushrooms (Amanita princeps).<ref>Template:Cite journal</ref>
Cultivation and usesEdit
Carpinus betulus is widely cultivated as an ornamental tree, for planting in gardens and parks throughout north west Europe. Because it stands up well to cutting back and has dense foliage, it has been much used in landscape gardening, mainly as tall hedges and for topiary. It was the classic tree used in French formal gardens for hedges in bosquets, as in the Gardens of Versailles, and in their English equivalent, the garden wilderness.Template:Citation needed
There are several cultivars, notably 'Fastigiata' or 'Pyramidalis', a fastigiate tree when young, which has become a popular urban street tree in the United Kingdom and other countries.Template:Citation needed 'Frans Fontaine' is a similar fastigiate cultivar.<ref>{{#invoke:citation/CS1|citation |CitationClass=web }}</ref> Both the species C. betulus and the cultivar 'Fastigiata' have gained the Royal Horticultural Society's Award of Garden Merit.<ref name=rhs>{{#invoke:citation/CS1|citation |CitationClass=web }}</ref><ref>{{#invoke:citation/CS1|citation |CitationClass=web }}</ref>
As early as Roman times, but also during the Thirty Years' War, defensive hedges (Landwehr) in Central Europe were largely planted from hornbeams. The bushes were chopped down with axes and broken down. The trees would then grow together with blackberries, dog roses, and other thorny bushes into impenetrable hedges. In the 11th century, for example, the Electorate of Mainz set up a large defensive forest called the Template:Ill. Many place names with the endings -hagen and -hain refer to such landwehrs.<ref name=laud>Template:Cite book</ref>
Hildegard of Bingen wrote of hornbeam being used as a plant in traditional medicine to treat vitiligo. The heated hornbeam chips were pressed onto the affected skin areas.<ref>Template:Cite book</ref> Hornbeam is used in Bach flower remedies as a treatment for exhaustion.<ref name=laud/>
The wood is heavy and hard, and is used for tools and building constructions. It also burns hot and slowly, making it very suitable for firewood.<ref>Hornbeam Tree - Gardening Central Template:Webarchive</ref> This was the reason for lopping and hence indirectly the saving of Epping Forest, where the hornbeam was a favoured pollarding tree.Template:Citation needed The wood has a very high calorific value of around 2,300 kWh/RM.<ref>{{#invoke:citation/CS1|citation |CitationClass=web }}</ref>
Notable examplesEdit
- The Last Tree - The only tree that survived the devastation of the Battle of Delville Wood in 1916. It is preserved as part of the Delville Wood South African National Memorial near Longueval.<ref>{{#invoke:citation/CS1|citation
|CitationClass=web }}</ref>
- Lincoln's Hornbeam - Planted by President Lincoln at the United States Botanic Garden in the Capitol grounds, Washington D.C.<ref>Template:Cite news</ref>
GalleryEdit
- Herchies JPG09.jpg
Tree in winter
- Carpinus betulus trunk.jpg
Old tree
- La Reid AR1aJPG.jpg
Hornbeam-covered walk
- Carpinus betulus MHNT.BOT.2004.0.494.jpg
Fruit and seeds
- Carpinus betulus bud.jpg
Leaf bud
- Carpinus betulus 'Fastigiata' Poundbury 20 P1310846.jpg
Newly planted Carpinus betulus 'Fastigiata' in Poundbury, Dorset, United Kingdom
- Common hornbeam growing in plastic tree shelter tube inside view.jpg
Common hornbeam in a tree protection tube planted 3Template:Nbspyears and 3Template:Nbspmonths earlier
ReferencesEdit
External linksEdit
- Den virtuella floran: Carpinus betulus distribution
- Carpinus betulus - information, genetic conservation units and related resources. European Forest Genetic Resources Programme (EUFORGEN)
