Edgar Adrian
Template:Short description Template:EngvarB Template:Use dmy dates {{#invoke:infobox|infoboxTemplate | bodyclass = vcard | bodystyle = {{#if:|width: {{{mainwidth}}}}} | child = {{{embed}}}
| abovestyle = font-size: 100%;
| above = {{#if:The Right Honourable|
}}
{{#if:Template:Postnominals|
}}
| subheaderstyle = font-size:125%; font-weight:bold;
| subheader = {{#ifeq:{{{embed}}}|yes||{{#if:|{{#if:|
}}}}}}
| image = {{#invoke:InfoboxImage|InfoboxImage|image=Edgar Douglas Adrian nobel.jpg|size=|sizedefault=frameless|upright=1|alt=black and white portrait photograph of Edgar Adrian, wearing a shirt, tie and jacket|suppressplaceholder=yes}} | image2 = {{#invoke:InfoboxImage|InfoboxImage|image=|size=|sizedefault=frameless|upright=1|alt=|suppressplaceholder=yes}} | image3 = {{#invoke:InfoboxImage|InfoboxImage|image=|sizedefault=frameless|upright=1|alt=|suppressplaceholder=yes}} | captionstyle = line-height:normal;padding-top:0.2em; | caption{{#if:|3|{{#if:|2}}}} = Adrian in 1932
| headerstyle = color: #202122; {{#ifeq:{{{embed}}}|yes|background:#eee|background:lavender}}
| data1 = {{#if:| {{#if:|| Template:Infobox officeholder/office}}}}Template:Infobox officeholder/office{{#if:|| Template:Infobox officeholder/office}}{{#if:|| {{#if:|| Template:Infobox officeholder/office}}}}{{#if:|| Template:Infobox officeholder/office}}{{#if:|| Template:Infobox officeholder/office}}{{#if:|| Template:Infobox officeholder/office}}{{#if:|| Template:Infobox officeholder/office}}{{#if:|| Template:Infobox officeholder/office}}{{#if:|| Template:Infobox officeholder/office}}{{#if:|| Template:Infobox officeholder/office}}{{#if:|| Template:Infobox officeholder/office}}{{#if:|| Template:Infobox officeholder/office}}{{#if:|| Template:Infobox officeholder/office}}{{#if:|| Template:Infobox officeholder/office}}{{#if:|| Template:Infobox officeholder/office}}{{#if:|| Template:Infobox officeholder/office}}{{#if:|| Template:Infobox officeholder/office}}
| data2 = | header3 = {{#if:Template:Birth dateHampstead, London, EnglandTemplate:Death date and ageCambridge, EnglandHester Adrian (m. 1923)Template:Plainlist|Personal details}} | label4 = Pronunciation | data4 =
| label5 = Born | data5 = {{#invoke:Separated entries|br
|1 = {{#if:|
}}
|2 = Template:Birth date |3 = Hampstead, London, England }}
| label6 = Died | data6 = {{#invoke:Separated entries|br|Template:Death date and age|Cambridge, England}}
| label7 = {{#ifexpr: Template:Strfind short
| Manner |{{#if:|Manner|Cause}} }} of death
| data7 = {{#if:||}}
| label8 = Resting place | class8 = label | data8 = {{#invoke:Separated entries|br||}}
| label9 = Citizenship | data9 =
| label10 = Nationality | data10 = {{#switch:{{#invoke:delink|delink|}} | {{#ifeq:Template:Country2nationality|{{#invoke:delink|delink|}}|{{#invoke:delink|delink|}}}} = | {{#ifeq:Template:Find country|England|British}} = | #default = }}
| label11 = Political party | data11 = {{#switch: | = | Democrat | Democratic | Democrat = Democratic | Republican | United States Republican Party | Republican | Republican Party = Republican | Conservative Party | Conservative = Conservative | Labour Party | Labour = Labour | Conservative Party | Conservative = Conservative | Liberal Party | Liberal = Liberal | KMT | Kuomintang | KMT | KMT | Kuomintang | Kuomintang (KMT) | Kuomintang (KMT) = Kuomintang | DPP | DPP | Democratic Progressive Party = Democratic Progressive Party | #default = }}
| label12 = Other political
affiliations
| data12 =
| label13 = Height | data13 = {{#if:|Template:Infobox person/height}}
| label14 = Spouse{{#if:|s|{{#invoke:Detect singular|pluralize|Hester Adrian (m. 1923)|likely=(s)|plural=s}}}} | data14 = Hester Adrian (m. 1923)
| label15 = Domestic partner{{#invoke:Detect singular|pluralize||likely=(s)|plural=s}} | data15 =
| label16 = Relations | data16 =
| label17 = Children | data17 = Template:Plainlist
| label18 = Parent{{#if:|{{#invoke:Detect singular|pluralize||likely=(s)|plural=s}}|{{#ifexpr:Template:Count > 1|s}}}} | data18 = {{#if:|{{{parents}}}|{{#invoke:list|unbulleted|{{#if:|{{{father}}} (father)}}|{{#if:|{{{mother}}} (mother)}}}}}}
| label19 = Relatives | data19 =
| label20 = Residence{{#invoke:Detect singular|pluralize||likely=(s)|plural=s}} | class20 = {{#if:Template:Death date and ageCambridge, England||label}} | data20 =
| label21 = Education | data21 =
| label22 = Alma mater | data22 =
| label23 = Occupation | data23 =
| label24 = Profession | data24 =
| label25 = Known for | data25 =
| label26 = Salary | data26 =
| label27 = Cabinet | data27 =
| label28 = Committees | data28 =
| label29 = Portfolio | data29 =
| label30 = {{#if:|Civilian awards|Awards}} | data30 =
| label31 = {{{blank1}}} | data31 =
| label32 = {{{blank2}}} | data32 =
| label33 = {{{blank3}}} | data33 =
| label34 = {{{blank4}}} | data34 =
| label35 = {{{blank5}}} | data35 =
| label36 = Signature | data36 = {{#if:|[[File:{{{signature}}}|{{#if:|{{{signature_size}}}|128x80px}}|class=skin-invert|alt=|Edgar Adrian's signature]]}}
| label37 = Website | data37 =
| label38 = Nickname{{#invoke:Detect singular|pluralize||likely=(s)|plural=s}} | data38 =
| header39 = {{#if:|Military service}}
| label40 = Allegiance | data40 =
| label41 = {{#if:||Branch/service}} | data41 =
| label42 = {{#if:||Years of service}} | data42 =
| label43 = {{#if:||Rank}} | data43 =
| label44 = {{#if:||Unit}} | data44 =
| label45 = Commands | data45 =
| label46 = {{#if:||Battles/wars}} | data46 =
| label47 = {{#if:|Military awards|Awards}} | data47 =
| label48 = {{{military_blank1}}} | data48 =
| label49 = {{{military_blank2}}} | data49 =
| label50 = {{{military_blank3}}} | data50 =
| label51 = {{{military_blank4}}} | data51 =
| label52 = {{{military_blank5}}} | data52 =
| data53 = Template:Infobox scientist | data54 = | data55 = | data56 = | data57 = | data58 = | belowstyle = border-top: 1px solid right;
| below =
{{#if:|Source: [{{{source}}}]}}
}}{{#if:|{{#ifeq:{{#ifeq:|no|yes}}|yes||}} }}{{#if:|{{#ifeq:{{#ifeq:|no|yes}}|yes||}} }}{{#if:|{{#if:||{{#ifeq:{{#ifeq:|no|yes}}|yes||}}}} }}{{#invoke:Check for unknown parameters|check|unknown=Template:Main other|preview=Page using Template:Infobox officeholder with unknown parameter "_VALUE_"|ignoreblank=y| regexp1 = 1blankname[%d]* | regexp2 = 1namedata[%d]* | regexp3 = 2blankname[%d]* | regexp4 = 2namedata[%d]* | regexp5 = 3blankname[%d]* | regexp6 = 3namedata[%d]* | regexp7 = 4blankname[%d]* | regexp8 = 4namedata[%d]* | regexp9 = 5blankname[%d]* | regexp10 = 5namedata[%d]* | allegiance | alma_mater | regexp11 = alongside[%d]* | alt | regexp12 = ambassador_from[%d]* | regexp13 = appointed[%d]* | regexp14 = appointer[%d]* | regexp15 = assembly[%d]* | awards | battles | battles_label | birth_date | birth_name | birth_place | birthname | regexp16 = blank[%d]* | bodyclass | branch | branch_label | cabinet | candidate | caption | categories | regexp17 = chancellor[%d]* | children | citizenship | regexp18 = co%-leader[%d]* | commands | committees | regexp19 = constituency[%d]* | regexp20 = constituency_AM[%d]* | regexp21 = constituency_MP[%d]* | regexp22 = convocation[%d]* | regexp23 = country[%d]* | regexp24 = data[%d]* | date | death_cause | death_date | death_manner | death_place | demo | regexp25 = deputy[%d]* | regexp26 = district[%d]* | education | election_date | embed | father | regexp28 = firstminister[%d]* | footnotes | regexp29 = governor[%d]* | regexp30 = governor_general[%d]* | regexp31 = governor%-general[%d]* | height | honorific_prefix | honorific-prefix | honorific_suffix | honorific-suffix | image | image name | image_name_alt | image_size | imagesize | image_upright | incumbent | regexp32 = jr/sr[%d]* | regexp33 = jr/sr and state[%d]* | known_for | regexp34 = leader[%d]* | regexp35 = legislature[%d]* | regexp36 = lieutenant[%d]* | regexp37 = lieutenant_governor[%d]* | mainwidth | regexp38 = majority[%d]* | regexp39 = majority_floor_leader[%d]* | regexp40 = majority_leader[%d]* | regexp41 = majorityleader[%d]* | mawards | regexp42 = military_blank[%d]* | regexp43 = military_data[%d]* | regexp44 = minister[%d]* | regexp45 = minister_from[%d]* | regexp46 = minority_floor_leader[%d]* | regexp47 = minority_leader[%d]* | regexp48 = minorityleader[%d]* | regexp49 = module[%d]* | regexp50 = monarch[%d]* | mother | name | nationality | native_name | native_name_lang | nickname | nocat | regexp51 = nominator[%d]* | nominee | occupation | regexp52 = office[%d]* | opponent | regexp53 = order[%d]* | otherparty | parents | regexp54 = parliament[%d]* | regexp55 = parliamentarygroup[%d]* | partner | party | party_election | portfolio | regexp56 = preceded[%d]* | regexp57 = preceding[%d]* | regexp58 = predecessor[%d]* | regexp59 = premier[%d]* | regexp60 = president[%d]* | regexp61 = primeminister[%d]* | regexp62 = prior_term[%d]* | profession | pronunciation | rank | rank_label | relations | relatives | residence | resting_place | resting_place_coordinates | restingplace | restingplacecoordinates | regexp63 = riding[%d]* | runningmate | salary | serviceyears | serviceyears_label | signature | signature_alt | signature_size | smallimage | smallimage_alt | source | speaker | speaker_office | spouse | spouses | regexp64 = state[%d]* | regexp65 = state_assembly[%d]* | regexp66 = state_delegate[%d]* | regexp67 = state_house[%d]* | regexp68 = state_legislature[%d]* | regexp69 = state_senate[%d]* | regexp70 = status[%d]* | regexp71 = suboffice[%d]* | regexp72 = subterm[%d]* | regexp73 = succeeded[%d]* | regexp74 = succeeding[%d]* | regexp75 = successor[%d]* | regexp76 = taoiseach[%d]* | regexp77 = term[%d]* | regexp78 = term_end[%d]* | regexp79 = term_label[%d]* | regexp80 = term_start[%d]* | regexp81 = termend[%d]* | regexp82 = termlabel[%d]* | regexp83 = termstart[%d]* | regexp84 = title[%d]* | unit | unit_label | regexp85 = vicegovernor[%d]* | regexp86 = vicepremier[%d]* | regexp87 = vicepresident[%d]* | regexp88 = viceprimeminister[%d]* | regexp89 = assuming[%d]* | website | width | year }}
Edgar Douglas Adrian, 1st Baron Adrian Template:Postnominals (30 November 1889 – 4 August 1977)<ref>GRO Register of Births: DEC 1889 1a 650 HAMPSTEAD – Edgar Douglas Adrian</ref><ref>GRO Register of Deaths: SEP 1977 9 0656 CAMBRIDGE – Edgar Douglas Adrian, DoB = 30 November 1889</ref> was an English electrophysiologist and recipient of the 1932 Nobel Prize for Physiology, won jointly with Sir Charles Sherrington for work on the function of neurons. He provided experimental evidence for the all-or-none law of nerves.<ref name="frs">Template:Cite journal</ref><ref name="Corsini2002">Template:Cite book</ref>
BiographyEdit
Adrian was born in Hampstead, London, the youngest son of Alfred Douglas Adrian, legal adviser to the Local Government Board, and Flora Lavinia Barton.<ref>Template:Cite book</ref>
He was educated at Westminster School and studied Natural Sciences at Trinity College, Cambridge, graduating in 1911. In 1913 he was elected to a fellowship of Trinity College on account of his research into the "all or none" law of nerves.
After completing a medical degree (MB BCh) in 1915, he undertook clinical work at St Bartholomew's Hospital London during World War I, treating soldiers with nerve damage and nervous disorders such as shell shock. Adrian returned to Cambridge as a lecturer gaining his doctorate (MD) in 1919 and in 1925 began research on the human sensory organs by electrical methods.
CareerEdit
Continuing earlier studies of Keith Lucas, he used a capillary electrometer and cathode-ray tube to amplify the signals produced by the nervous system and was able to record the electrical discharge of single nerve fibres under physical stimulus. (It seems he used frogs in his experiments<ref>{{#invoke:citation/CS1|citation |CitationClass=web }}</ref>) An accidental discovery by Adrian in 1928 proved the presence of electricity within nerve cells. Adrian said,
I had arranged electrodes on the optic nerve of a toad in connection with some experiments on the retina. The room was nearly dark and I was puzzled to hear repeated noises in the loudspeaker attached to the amplifier, noises indicating that a great deal of impulse activity was going on. It was not until I compared the noises with my own movements around the room that I realised I was in the field of vision of the toad's eye and that it was signalling what I was doing.
A key result, published in 1928, stated that the excitation of the skin under constant stimulus is initially strong but gradually decreases over time, whereas the sensory impulses passing along the nerves from the point of contact are constant in strength, yet are reduced in frequency over time, and the sensation in the brain diminishes as a result.
Extending these results to the study of pain causes by the stimulus of the nervous system, he made discoveries about the reception of such signals in the brain and spatial distribution of the sensory areas of the cerebral cortex in different animals. These conclusions lead to the idea of a sensory map, called the homunculus, in the somatosensory system.
Later, Adrian used the electroencephalogram to study the electrical activity of the brain in humans. His work on the abnormalities of the Berger rhythm paved the way for subsequent investigation in epilepsy and other cerebral pathologies. He spent the last portion of his research career investigating olfaction.
Positions that he held during his career included Foulerton Professor 1929–1937; Professor of Physiology in the University of Cambridge 1937–1951; President of the Royal Society 1950–1955; Master of Trinity College, Cambridge, 1951–1965; president of the Royal Society of Medicine 1960–1962; Chancellor of the University of Cambridge 1967–1975; Chancellor of the University of Leicester 1957–1971. Adrian was elected a Foreign Honorary Member of the American Academy of Arts and Sciences and the American Philosophical Society in 1938.<ref name=AAAS>{{#invoke:citation/CS1|citation |CitationClass=web }}</ref><ref>{{#invoke:citation/CS1|citation |CitationClass=web }}</ref> He was elected an International Member of the United States National Academy of Sciences in 1941.<ref>{{#invoke:citation/CS1|citation |CitationClass=web }}</ref> In 1946 he became foreign member of the Royal Netherlands Academy of Arts and Sciences.<ref>{{#invoke:citation/CS1|citation |CitationClass=web }}</ref> In 1942 he was awarded the Order of Merit and in the 1955 New Year Honours was created Baron Adrian of Cambridge in the County of Cambridge.<ref>UK list: Template:London Gazette</ref><ref>{{#invoke:citation/CS1|citation |CitationClass=web }}</ref>
FamilyEdit
On 14 June 1923 Edgar Adrian married Hester Agnes Pinsent, who was the daughter of Ellen Pinsent and sister of David Pinsent. Together they had three children, first a daughter and then twins:
- Anne Pinsent Adrian, who married the physiologist Richard Keynes, who was the great-grandson of Charles Darwin. The couple had four children.
- Adrian Keynes (1946–1974)
- Randal Keynes (*1948)
- Roger Keynes (*1951)
- Simon Keynes (*1952)
- Richard Adrian, 2nd Baron Adrian (1927–1995)
- The couple had no children.
- Jennet Adrian (b. 1927), who married Peter Watson Campbell
- Further family details unknown.<ref>Peter Townend, ed., Burke's Peerage and Baronetage, 105th edition (London, U.K.: Burke's Peerage Ltd, 1970), page 27.</ref><ref>Template:Cite journal</ref>
BibliographyEdit
- The Basis of Sensation (1928)
- The Mechanism of Nervous Action (1932)
- Factors Determining Human Behavior (1937)
ArmsEdit
See alsoEdit
ReferencesEdit
External linksEdit
- Template:Nobelprize including the Nobel Lecture on 12 December 1932 The Activity of the Nerve Fibres
- The Master of Trinity at Trinity College, Cambridge
Template:S-start Template:S-aca Template:S-bef Template:S-ttl Template:S-aft Template:S-break Template:S-new Template:S-ttl Template:S-aft Template:S-break Template:S-bef Template:S-ttl Template:S-aft Template:S-break Template:S-npo Template:S-bef Template:S-ttl Template:S-aft Template:S-break Template:S-reg Template:S-new Template:S-ttl Template:S-aft Template:S-end
Template:Copley Medallists 1901-1950 Template:Nobel Prize in Physiology or Medicine Laureates 1926-1950 Template:1932 Nobel Prize winners Template:Royal Society presidents 1900s Template:Presidents of the Royal Society of Medicine Template:Masters of Trinity College, Cambridge Template:Chancellors of the University of Cambridge Template:Authority control