Hepatica

File:Hepatica transsylvanica.jpg

Hepatica transsilvanica

Hepatica (hepatica,<ref>Gleason, H.A. 1952. The new Britton and Brown Illustrated flora of the Northeastern United States and adjacent Canada. Volume 2: The chloripetalous Dicotyledoneae. Hafner Press, New York.</ref> liverleaf,<ref>Bailey, L.H.; Bailey, E.Z.; the staff of the Liberty Hyde Bailey Hortorium. 1976. Hortus third: A concise dictionary of plants cultivated in the United States and Canada. Macmillan, New York.</ref> or liverwort)<ref>Webster's Third International Dictionary</ref> is a genus of herbaceous perennials in the buttercup family, native to central and northern Europe, Asia and eastern North America. Some botanists include Hepatica within a wider interpretation of Anemone.<ref name="Lloyd">{{#invoke:citation/CS1|citation |CitationClass=web }}</ref><ref>Template:Cite journal</ref>

Description

Bisexual flowers with pink, purple, blue, or white sepals and three green bracts appear singly on hairy stems from late winter to spring. Butterflies, moths, bees, flies and beetles are known pollinators.

The leaves are basal, leathery, and usually three-lobed, remaining over winter.

Taxonomy

Hepatica was described by the English botanist Philip Miller in 1754.Template:R It was proposed as a subgenus of Anemone in 1836,Template:R but later segregated into genus Hepatica.

Taxa

File:HepaticaNobilisMacro.jpg

Hepatica nobilis flowers

File:Hepatica in Lithuania.jpg

Hepatica in Europos Parkas, Lithuania

File:Hepatica nobilis var. pyrenaica .jpg

Hepatica nobilis in Aínsa, Spain

Template:As of, Kew's Plants of the World Online (POWO) accepts 7 species and one hybrid in the genus Hepatica:Template:R

One infraspecific taxon is also recognized by POWO:Template:R

• Hepatica nobilis var. japonica Template:Small
  • Synonym: Hepatica asiatica Template:Small
  • Synonym: Hepatica insularis Template:Small

Hepatica can be divided into two series with respect to leaf edge:

Series Triloba

The leaves of the series Triloba Ulbr.<ref name="Ulbr">Ulbrich, O.E.: Über die systematische Gliederung und geographische Verbreitung der Gattung Anemone L. Bot. Jahrb. Syst. (1905) 37: 172 - 257, 38: 257 - 334.</ref> Tamura:<ref name=Tamu>Tamura, M.: Morphology, ecology and phylogeny of the Ranunculaceae” VII. Science reports of South College, North College of Osaka University, Japan 16:21-43, 1968.</ref> are three-lobed with a smooth leaf edge.

• Hepatica acutiloba: Eastern Canada, Midwestern United States, Eastern United StatesTemplate:R
• Hepatica americana: Central Canada, Eastern Canada, Midwestern United States, Eastern United StatesTemplate:R
• Hepatica maxima: endemic to Ulleung-do island (South Korea)Template:R
• Hepatica nobilis:
  • Hepatica nobilis var. nobilis: Europe to Western SiberiaTemplate:R
  • Hepatica nobilis var. japonica: Russian Far East, North China, Central China, East China, Korea, and Japan (Honshu, Shikoku, north of Kyushu island)Template:R

Series Angulosa

The leaves of series Angulosa (Ulbr.)<ref name=Ulbr /> Tamura<ref name= Tamu /> are three- to five-lobed with a crenate leaf edge.

• Hepatica falconeri — Mountain forests of Central Asia; India: Northwest Himalayas (Himachal Pradesh, Jammu and Kashmir); Northwest China (Tienshan); Kyrgyzstan and Tajikistan (Pamir-Alai); North Pakistan, Kazakhstan (North Tienshan)Template:R
• Hepatica henryi: North Central China, South Central China (western Hubei, northern Hunan, Sichuan, Shaanxi)Template:R<ref>STEWARD, A.N.: in Rhodora 29: 53. 1927</ref><ref>Peters, Jürgen: Das etwas andere Leberblümchen: Hepatica yamatutai Nakai in ‚Gartenbotanische Blätter‘ 5/2000 der Gartenbotanischen Vereinigung in Deutschland</ref>
• Hepatica × media: RomaniaTemplate:R
• Hepatica transsilvanica: Romania (Carpathian Mountains, Transylvania)Template:R

Etymology

The word hepatica derives from the Greek {{#invoke:Lang|lang}} Template:Grc-tr, from Template:Wikt-lang Template:Grc-tr 'liver', because its three-lobed leaf was thought to resemble the human liver.<ref name=SOED>Template:Cite book</ref>

Distribution

Plants of genus Hepatica are native to Europe, Asia, and North America.Template:R

• Europe: Albania, Austria, the Baltic states, Belarus, Bulgaria, Corsica, Czechoslovakia, Denmark, European Russia, Finland, France, Germany, Greece, Hungary, Italy, Norway, Poland, Romania, Spain, Sweden, Switzerland, Ukraine, Yugoslavia
• Central Asia: Kazakhstan, Kyrgyzstan, Tajikistan, Western Siberia
• Eastern Asia: North China, South Central China, East China, Japan, Korea, Manchuria, Primorsky Krai
• South Asia: Pakistan, Western Himalaya
• Canada: Manitoba, New Brunswick, Nova Scotia, Ontario, Québec
• United States: Alabama, Arkansas, Connecticut, Delaware, District of Columbia, Florida, Georgia, Illinois, Indiana, Iowa, Kentucky, Maine, Maryland, Massachusetts, Michigan, Minnesota, Mississippi, Missouri, New Hampshire, New Jersey, New York, North Carolina, Ohio, Pennsylvania, Rhode Island, South Carolina, Tennessee, Vermont, Virginia, West Virginia, Wisconsin

Plants of the genus have been introduced to Belgium.Template:R

Cultivation

Hepatica cultivation has been popular in Japan since the 18th century (mid-Edo period), where flowers with doubled petals and a range of colour patterns have been developed.<ref>Template:Cite journal</ref>

Noted for its tolerance of alkaline limestone-derived soils, Hepatica may grow in a wide range of conditions; it can be found either in deeply shaded deciduous (especially beech) woodland and scrub or grassland in full sun. Hepatica will also grow in both sandy and clay-rich substrates, being associated with limestone. Moist soil and winter snowfall are required; Hepatica is tolerant of winter snow cover, but less so of dry frost.

Propagation is done by seeds or by dividing vigorous clumps in spring. However, seedlings take several years to reach bloom size, and divided plants are slow to thicken.

Uses

Hepatica was once used as a medicinal herb. Owing to the doctrine of signatures, the plant was once thought to be an effective treatment for liver disorders. Although poisonous in large doses, the leaves and flowers may be used as an astringent, as a demulcent for slow-healing injuries, and as a diuretic.<ref name="Lloyd"/>

External links

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