The tree line is the edge of a habitat at which trees are capable of growing and beyond which they are not. It is found at high elevations and high latitudes. Beyond the tree line, trees cannot tolerate the environmental conditions (usually low temperatures, extreme snowpack, or associated lack of available moisture).<ref name="taiga">Template:Cite book</ref>Template:Rp The tree line is sometimes distinguished from a lower timberline, which is the line below which trees form a forest with a closed canopy.<ref name="ecosystem">Template:Cite book</ref>Template:Rp<ref>Template:Cite book</ref>Template:Rp
At the tree line, tree growth is often sparse, stunted, and deformed by wind and cold. This is sometimes known as {{#invoke:Lang|lang}} (German for "crooked wood").<ref name="landabove">Template:Cite book</ref>Template:Rp
The tree line often appears well-defined, but it can be a more gradual transition. Trees grow shorter and often at lower densities as they approach the tree line, above which they are unable to grow at all.<ref name=landabove/>Template:Rp Given a certain latitude, the tree line is approximately 300 to 1000 meters below the permanent snow line and roughly parallel to it.<ref>{{#invoke:citation/CS1|citation |CitationClass=web }}</ref>
CausesEdit
Due to their vertical structure, trees are more susceptible to cold than more ground-hugging forms of plants.<ref name=coldtreeline/> Summer warmth generally sets the limit to which tree growth can occur: while tree line conifers are very frost-hardy during most of the year, they become sensitive to just 1 or 2 degrees of frost in mid-summer.<ref name="tran">Template:Cite book</ref><ref>Template:Cite book</ref> A series of warm summers in the 1940s seems to have permitted the establishment of "significant numbers" of spruce seedlings above the previous treeline in the hills near Fairbanks, Alaska.<ref name="vier4">Template:Cite journal</ref><ref name="vier8">Template:Cite book</ref> Survival depends on a sufficiency of new growth to support the tree. Wind can mechanically damage tree tissues directly, including blasting with windborne particles, and may also contribute to the desiccation of foliage, especially of shoots that project above the snow cover.Template:Cn
The actual tree line is set by the mean temperature, while the realized tree line may be affected by disturbances, such as logging,<ref name="coldtreeline">Template:Cite journal</ref> or grazing<ref>Template:Cite journal</ref> Most human activities cannot change the actual tree line, unless they affect the climate.<ref name=coldtreeline/> The tree line follows the line where the seasonal mean temperature is approximately Template:Convert.<ref>Template:Cite journal</ref><ref name=coldtreeline/> The seasonal mean temperature is taken over all days whose mean temperature is above Template:Convert. A growing season of 94 days above that temperature is required for tree growth.<ref>Template:Cite journal</ref>
Because of climate change, which leads to earlier snow melt and favorable conditions for tree establishment, the tree line in North Cascades National Park has risen more than Template:Convert in 50 years.<ref>{{#invoke:citation/CS1|citation |CitationClass=web }}</ref>
TypesEdit
Several types of tree lines are defined in ecology and geography:
AlpineEdit
An alpine tree line is the highest elevation that sustains trees; higher up it is too cold, or the snow cover lasts for too much of the year, to sustain trees.<ref name=ecosystem/>Template:Rp The climate above the tree line of mountains is called an alpine climate,<ref name="plantlife">Template:Cite book</ref>Template:Rp and the habitat can be described as the alpine zone.<ref name="romo">{{#invoke:citation/CS1|citation |CitationClass=web }}</ref> Treelines on north-facing slopes in the northern hemisphere are lower than on south-facing slopes, because the increased shade on north-facing slopes means the snowpack takes longer to melt. This shortens the growing season for trees.<ref name="rockymtns">Template:Cite book</ref>Template:Rp In the southern hemisphere, the south-facing slopes have the shorter growing season.
The alpine tree line boundary is seldom abrupt: it usually forms a transition zone between closed forest below and treeless alpine zone above. This zone of transition occurs "near the top of the tallest peaks in the northeastern United States, high up on the giant volcanoes in central Mexico, and on mountains in each of the 11 western states and throughout much of Canada and Alaska".<ref name="arno" /> Environmentally dwarfed shrubs (krummholz) commonly form the upper limit.
The decrease in air temperature with increasing elevation creates the alpine climate. The rate of decrease can vary in different mountain chains, from Template:Convert per Template:Convert of elevation gain in the dry mountains of the western United States,<ref name="arno" /> to Template:Convert per Template:Convert in the moister mountains of the eastern United States.<ref name="baker">Template:Cite journal</ref> Skin effects and topography can create microclimates that alter the general cooling trend.<ref name="gei">Template:Cite book</ref>
Compared with arctic tree lines, alpine tree lines may receive fewer than half of the number of degree days (above Template:Convert) based on air temperature, but because solar radiation intensities are greater at alpine than at arctic tree lines the number of degree days calculated from leaf temperatures may be very similar.<ref name="arno" />
At the alpine tree line, tree growth is inhibited when excessive snow lingers and shortens the growing season to the point where new growth would not have time to harden before the onset of fall frost. Moderate snowpack, however, may promote tree growth by insulating the trees from extreme cold during the winter, curtailing water loss,<ref name="sowell">Template:Cite journal</ref> and prolonging a supply of moisture through the early part of the growing season. However, snow accumulation in sheltered gullies in the Selkirk Mountains of southeastern British Columbia causes the tree line to be Template:Convert lower than on exposed intervening shoulders.<ref name="shaw">Template:Cite journal</ref>
In some mountainous areas, higher elevations above the condensation line, or on equator-facing and leeward slopes, can result in low rainfall and increased exposure to solar radiation. This dries out the soil, resulting in a localized arid environment unsuitable for trees. Many south-facing ridges of the mountains of the Western U.S. have a lower treeline than the northern faces because of increased sun exposure and aridity. Hawaii's treeline of about Template:Convert is also above the condensation zone and results due to a lack of moisture.Template:Cn
ExposureEdit
On coasts and isolated mountains, the tree line is often much lower than in corresponding altitudes inland and in larger, more complex mountain systems, because strong winds reduce tree growth. In addition, the lack of suitable soil, such as along talus slopes or exposed rock formations, prevents trees from gaining an adequate foothold and exposes them to drought and sun.Template:Cn
ArcticEdit
The Arctic tree line is the northernmost latitude in the Northern Hemisphere where trees can grow; farther north, it is too cold all year round to sustain trees.<ref>Template:Cite book</ref> Extremely low temperatures, especially when prolonged, can freeze the internal sap of trees, killing them. In addition, permafrost in the soil can prevent trees from getting their roots deep enough for the necessary structural support.Template:Cn
Unlike alpine tree lines, the northern tree line occurs at low elevations. The Arctic forest–tundra transition zone in northwestern Canada varies in width, perhaps averaging Template:Convert and widening markedly from west to east,<ref name="timoney">Template:Cite journal</ref> in contrast with the telescoped alpine timberlines.<ref name="arno">Template:Cite book</ref> North of the arctic tree line lies the low-growing tundra, and southwards lies the boreal forest.
Two zones can be distinguished in the Arctic tree line:<ref name="love">Template:Cite journal</ref><ref name="hare2">Template:Cite journal</ref> a forest–tundra zone of scattered patches of krummholz or stunted trees, with larger trees along rivers and on sheltered sites set in a matrix of tundra; and "open boreal forest" or "lichen woodland", consisting of open groves of erect trees underlain by a carpet of Cladonia spp. lichens.<ref name="love" /> The proportion of trees to lichen mat increases southwards towards the "forest line", where trees cover 50 percent or more of the landscape.<ref name="arno" /><ref name="black">Template:Cite journal</ref>
AntarcticEdit
A southern treeline exists in the New Zealand Subantarctic Islands and the Australian Macquarie Island, with places where mean annual temperatures above Template:Convert support trees and woody plants, and those below Template:Convert do not.<ref>Template:WWF ecoregion</ref> Another treeline exists in the southwesternmost parts of the Magellanic subpolar forests ecoregion, where the forest merges into the subantarctic tundra (termed Magellanic moorland or Magellanic tundra).<ref>Template:WWF ecoregion</ref> For example, the northern halves of Hoste and Navarino Islands have Nothofagus antarctica forests but the southern parts consist of moorlands and tundra.
Tree species near tree lineEdit
Some typical Arctic and alpine tree line tree species (note the predominance of conifers):
AustraliaEdit
- Snow gum (Eucalyptus pauciflora)
EurasiaEdit
- Dahurian larch (Larix gmelinii)
- Macedonian pine (Pinus peuce)
- Swiss pine (Pinus cembra)
- Mountain pine (Pinus mugo)
- Arctic white birch (Betula pubescens subsp. tortuosa)
- Rowan<ref>Template:Cite book</ref> (Sorbus aucuparia)
North AmericaEdit
- Subalpine fir (Abies lasiocarpa)<ref name=rockymtns/>Template:Rp
- Subalpine larch (Larix lyallii)<ref name="cancyc">Template:Cite encyclopedia</ref>
- Mountain hemlock (Tsuga mertensiana)
- Alaska yellow cedar (Chaemaecyparis nootkatensis)
- Engelmann spruce (Picea engelmannii)<ref name=rockymtns/>Template:Rp
- Whitebark pine (Pinus albicaulis)<ref name=cancyc/>
- Great Basin bristlecone pine (Pinus longaeva)
- Rocky Mountains bristlecone pine (Pinus aristata)
- Foxtail pine (Pinus balfouriana)
- Limber pine (Pinus flexilis)
- Potosi pinyon (Pinus culminicola)
- Black spruce (Picea mariana)<ref name=taiga/>Template:Rp
- White spruce (Picea glauca)
- Tamarack (Larix laricina)
- Hartweg's pine (Pinus hartwegii)
South AmericaEdit
- Antarctic beech (Nothofagus antarctica)
- Lenga beech (Nothofagus pumilio)<ref name="Fajardo">Template:Cite journal</ref>
- Alder (Alnus acuminata)
- Pino del cerro (Podocarpus parlatorei)
- Polylepis (Polylepis tarapacana)
- Eucalyptus (not native to South America but grown in large amounts in the high Andes).<ref>Template:Cite journal</ref>
Worldwide distributionEdit
Alpine tree linesEdit
The alpine tree line at a location is dependent on local variables, such as aspect of slope, rain shadow and proximity to either geographical pole. In addition, in some tropical or island localities, the lack of biogeographical access to species that have evolved in a subalpine environment can result in lower tree lines than one might expect by climate alone.Template:Cn
Averaging over many locations and local microclimates, the treeline rises Template:Convert when moving 1 degree south from 70 to 50°N, and Template:Convert per degree from 50 to 30°N. Between 30°N and 20°S, the treeline is roughly constant, between Template:Convert.<ref name=korner/>
Here is a list of approximate tree lines from locations around the globe: Template:Clear right
| Location | Approx. latitude | Approx. elevation of tree line | Notes | |
|---|---|---|---|---|
| (m) | (ft) | |||
| Finnmarksvidda, Norway | 69°N | Template:Convert | At 71°N, near the coast, the tree-line is below sea level (Arctic tree line). | |
| Abisko, Sweden | 68°N | Template:Convert | <ref name=korner/> | |
| Chugach Mountains, Alaska | 61°N | Template:Convert | Tree line around Template:Convert or lower in coastal areas | |
| Southern Norway | 61°N | Template:Convert | Much lower near the coast, down to Template:Convert. | |
| Scotland, United Kingdom | 57°N | Template:Convert | citation | CitationClass=web
}}</ref>Template:Rp |
| Northern Quebec | 56°N | Template:Convert | The cold Labrador Current originating in the arctic makes eastern Canada the sea-level region with the most southern tree-line in the northern hemisphere. | |
| Southern Urals | 55°N | Template:Convert | ||
| Canadian Rockies | 51°N | Template:Convert | ||
| Tatra Mountains | 49°N | Template:Convert | ||
| Olympic Mountains, Washington, United States | 47°N | Template:Convert | Heavy winter snowpack buries young trees until late summer | |
| Swiss Alps | 47°N | Template:Convert | <ref name=korner1/> | |
| Mount Katahdin, Maine, United States | 46°N | Template:Convert | ||
| Eastern Alps, Austria, Italy | 46°N | Template:Convert | More exposure to cold Russian winds than Western Alps | |
| Sikhote-Alin, Russia | 46°N | Template:Convert | citation | CitationClass=web
}}</ref> |
| Alps of Piedmont, Northwestern Italy | 45°N | Template:Convert | ||
| New Hampshire, United States | 44°N | Template:Convert | citation | CitationClass=web
}}</ref> Some peaks have even lower treelines because of fire and subsequent loss of soil, such as Grand Monadnock and Mount Chocorua. |
| Wyoming, United States | 43°N | Template:Convert | ||
| Caucasus Mountains | 42°N | Template:Convert | citation | CitationClass=web
}}</ref> |
| Rila and Pirin Mountains, Bulgaria | 42°N | Template:Convert | Up to Template:Convert on favorable locations. Mountain Pine is the most common tree line species. | |
| Pyrenees Spain, France, Andorra | 42°N | Template:Convert | Mountain Pine is the tree line species | |
| Steens Mountain, Oregon, US | 42°N | Template:Convert | ||
| Wasatch Mountains, Utah, United States | 40°N | Template:Convert | Higher (nearly Template:Convert in the Uintas) | |
| Rocky Mountain NP, CO, United States | 40°N | Template:Convert | <ref name=korner/> On warm southwest slopes | |
| Template:Convert | On northeast slopes | |||
| Yosemite, CA, United States | 38°N | Template:Convert | <ref name="Schoenherr">Template:Cite book</ref> West side of Sierra Nevada | |
| Template:Convert | <ref name=Schoenherr/> East side of Sierra Nevada | |||
| Sierra Nevada, Spain | 37°N | Template:Convert | Precipitation low in summer | |
| Japanese Alps | 36°N | Template:Convert | ||
| Khumbu, Himalaya | 28°N | Template:Convert | <ref name=korner/> | |
| Yushan, Taiwan | 23°N | Template:Convert | citation | CitationClass=web
}}</ref> Strong winds and poor soil restrict further grow of trees. |
| Hawaii, United States | 20°N | Template:Convert | <ref name=korner/> Geographic isolation and no local tree species with high tolerance to cold temperatures. | |
| Pico de Orizaba, Mexico | 19°N | Template:Convert | citation | CitationClass=web
}}</ref> |
| Costa Rica | 9.5°N | Template:Convert | ||
| Mount Kinabalu, Borneo | 6.1°N | Template:Convert | citation | CitationClass=web
}}</ref> |
| Mount Kilimanjaro, Tanzania | 3°S | Template:Convert | <ref name="korner">Template:Cite journal</ref> Upper limit of forest trees; woody ericaeous scrub grows up to 3900m | |
| New Guinea | 6°S | Template:Convert | <ref name=korner/> | |
| Andes, Peru | 11°S | Template:Convert | East side; on west side tree growth is restricted by dryness | |
| Andes, Bolivia | 18°S | Template:Convert | Western Cordillera; highest treeline in the world on the slopes of Sajama Volcano (Polylepis tarapacana) | |
| Template:Convert | Eastern Cordillera; treeline is lower because of lower solar radiation (more humid climate) | |||
| Sierra de Córdoba, Argentina | 31°S | Template:Convert | Precipitation low above trade winds, also high exposure | |
| Australian Alps, New South Wales, Australia | 36°S | |||
| Template:Convert | citation | CitationClass=web
}}</ref> | ||
| Andes, Laguna del Laja, Chile | 37°S | Template:Convert | Temperature rather than precipitation restricts tree growth<ref name="Lara">Template:Cite journal</ref> | |
| Mount Taranaki, North Island, New Zealand | 39°S | Template:Convert | Strong maritime influence serves to cool summer and restrict tree growth | |
| Northeast Tasmania, Australia | 41°S | Template:Convert | Although sheltered on the leeward side of the island, summers are still cool for the latitude. | |
| Southwest Tasmania, Australia | 43°S | Template:Convert | Exposed to the westerly storm track, summer is extraordinarily cool for the latitude, with frequent summer snow. Springtime receives an extreme amount of cold, heavy precipitation; winds are likewise extreme. | |
| Fiordland, South Island, New Zealand | 45°S | Template:Convert | Very snowy springs, strong cold winds and cool summers with frequent summer snow restrict tree growthTemplate:Citation needed | |
| Lago Argentino, Argentina | 50°S | Template:Convert | Nothofagus pumilio<ref>Template:Cite journal</ref> | |
| Torres del Paine, Chile | 51°S | Template:Convert | Strong influence from the Southern Patagonian Ice Field serves to cool summer and restrict tree growth<ref name="Patagonia">Template:Cite journal</ref> | |
| Navarino Island, Chile | 55°S | Template:Convert | Strong maritime influence serves to cool summer and restrict tree growth<ref name=Patagonia/> | |
Arctic tree linesEdit
Like the alpine tree lines shown above, polar tree lines are heavily influenced by local variables such as aspect of slope and degree of shelter. In addition, permafrost has a major impact on the ability of trees to place roots into the ground. When roots are too shallow, trees are susceptible to windthrow and erosion. Trees can often grow in river valleys at latitudes where they could not grow on a more exposed site. Maritime influences such as ocean currents also play a major role in determining how far from the equator trees can grow as well as the warm summers experienced in extreme continental climates.Template:Cn In northern inland Scandinavia there is substantial maritime influence on high parallels that keep winters relatively mild, but enough inland effect to have summers well above the threshold for the tree line. Here are some typical polar treelines:
| Location | Approx. longitude | Approx. latitude of tree line | Notes |
|---|---|---|---|
| Norway | Template:Sort | 70°N | The North Atlantic current makes Arctic climates in this region warmer than other coastal locations at comparable latitude. In particular the mildness of winters prevents permafrost. |
| West Siberian Plain | Template:Sort | 66°N | |
| Central Siberian Plateau | Template:Sort | 72°N | Extreme continental climate means the summer is warm enough to allow tree growth at higher latitudes, extending to northernmost forests of the world at 72°28'N at Ary-Mas (102° 15' E) in the Novaya River valley, a tributary of the Khatanga River and the more northern Lukunsky grove at 72°31'N, 105° 03' E east from Khatanga River. |
| Russian Far East (Kamchatka and Chukotka) | Template:Sort | 60°N | The Oyashio Current and strong winds affect summer temperatures to prevent tree growth. The Aleutian Islands are almost completely treeless. |
| Alaska, United States | Template:Sort | 68°N | Trees grow north to the south-facing slopes of the Brooks Range. The mountains block cold air coming off of the Arctic Ocean. |
| Northwest Territories, Canada | Template:Sort | 69°N | Reaches north of the Arctic Circle because of the continental nature of the climate and warmer summer temperatures. |
| Nunavut | Template:Sort | 61°N | Influence of the very cold Hudson Bay moves the treeline southwards. |
| Labrador Peninsula | Template:Sort | 56°N | Very strong influence of the Labrador Current on summer temperatures as well as altitude effects (much of Labrador is a plateau). In parts of Labrador, the treeline extends as far south as 53°N. Along the coast the northernmost trees are at 58°N in Napartok Bay. |
| Greenland | Template:Sort | 69°N | Determined by experimental tree planting in the absence of native trees because of isolation from natural seed sources; a very few trees are surviving, but growing slowly, at Søndre Strømfjord, 67°N. There is one natural forest in the Qinngua Valley. |
Antarctic tree linesEdit
Trees exist on Tierra del Fuego (55°S) at the southern end of South America, but generally not on subantarctic islands and not in Antarctica. Therefore, there is no explicit Antarctic tree line.Template:Cn
Kerguelen Island (49°S), South Georgia (54°S), and other subantarctic islands are all so heavily wind-exposed and with a too-cold summer climate (tundra) that none have any indigenous tree species. The Falkland Islands (51°S) summer temperature is near the limit, but the islands are also treeless, although some planted trees exist.Template:Cn
Antarctic Peninsula is the northernmost point in Antarctica (63°S) and has the mildest weather—it is located Template:Convert from Cape Horn on Tierra del Fuego—yet no trees survive there; only a few mosses, lichens, and species of grass do so. In addition, no trees survive on any of the subantarctic islands near the peninsula.Template:Cn
Southern Rata forests exist on Enderby Island and Auckland Islands (both 50°S) and these grow up to an elevation of Template:Convert in sheltered valleys. These trees seldom grow above Template:Convert in height and they get smaller as one gains altitude, so that by Template:Convert they are waist-high. These islands have only between 600 and 800 hours of sun annually. Campbell Island (52°S) further south is treeless, except for one stunted Spruce, probably planted in 1907.<ref>Template:Cite news</ref> The climate on these islands is not severe, but tree growth is limited by almost continual rain and wind. Summers are very cold with an average January temperature of Template:Convert. Winters are mild Template:Convert but wet. Macquarie Island (Australia) is located at 54°S and has no vegetation beyond snow grass and alpine grasses and mosses.Template:Citation needed
See alsoEdit
- Template:Portal-inline
- Montane ecosystems
- Ecotone: a transition between two adjacent ecological communities
- Edge effects: the effect of contrasting environments on an ecosystem
- Massenerhebung effect
- Snow line