Template:Short description Template:About Template:Automatic taxobox
Saxifraga is the largest genus in the family Saxifragaceae, containing about 473 species of holarctic perennial plants, known as saxifrages<ref name = powo/><ref name="Cam">{{#invoke:citation/CS1|citation |CitationClass=web }}</ref> or rockfoils.<ref>Roger Spencer, ed. Horticultural Flora of South-Eastern Australia. UNSW Press, 2002. p. 81. Template:ISBN</ref> The Latin word saxifraga means literally "stone-breaker", from Latin {{#invoke:Lang|lang}} ("rock" or "stone") + {{#invoke:Lang|lang}} ("to break"). It is usually thought to indicate a medicinal use for treatment of urinary calculi (known as kidney or bladder stones), rather than breaking rocks apart.<ref name="Cam"/><ref>Template:Cite book</ref>
DescriptionEdit
Most saxifrages are small perennial, biennial (e.g. S. adscendens) or annual (e.g. S. tridactylites) herbaceous plants whose basal or cauline leaves grow close to the ground, often in a rosette. The leaves typically have a more or less incised margin; they may be succulent, needle-like and/or hairy, reducing evaporation.Template:SfnTemplate:Sfn<ref name="RHSAZ"/>
The inflorescence or single flower clusters rise above the main plant body on naked stalks. The small actinomorphic hermaphrodite flowers have five petals and sepals and are usually white, but red to yellow in some species. Stamens, usually 10, rarely 8, insert at the junction of the floral tube and ovary wall, with filaments subulate or clavate. As in other primitive eudicots, some of the 5 or 10 stamens may appear petal-like.Template:Citation needed It lives in tundral ecosystems.Template:SfnTemplate:SfnTemplate:Sfn
TaxonomyEdit
A genus of about 473 species.<ref name = powo/> The former monotypic genus Saxifragella has been submersed within Saxifraga, the largest genus in Saxifragaceae, as Saxifraga bicuspidata.Template:SfnTemplate:Sfn Also, the genus Saxifragopsis (strawberry saxifrage) was previously included in Saxifraga.<ref name = usda2005>{{#invoke:citation/CS1|citation |CitationClass=web }}</ref>
SubdivisionEdit
Based on morphological criteria, up to 15 sections were recognised.Template:Sfn Subsequent molecular phylogenetic studies reduced this to 13 sections with 9 subsections. The former sections Micranthes and Merkianae are more closely related to the Boykinia and Heuchera clades.Template:Sfn Modern floras separate these groups as the genus Micranthes.<ref>Flora of China</ref>Template:Sfn
The thirteen sections (with subsections) are:Template:Sfn
- Irregulares
- Saxifragella
- Pseudocymbalaria
- Bronchiales
- Ciliatae
- Cymbalaria
- Cotylea
- Gymnopera
- Mesogyne
- Trachyphyllum
- Ligulatae
- Porphyrion
- Squarrosae
- Mutatae
- Oppositifoliae
- Florulentae
- Kabschia
- Saxifraga
- Tridactylites
- Androsaceae
- Arachnoideae
- Saxifraga
Selected speciesEdit
{{#invoke:Labelled list hatnote|labelledList|Main article|Main articles|Main page|Main pages}} Template:Div col
- Saxifraga adscendens – ascending saxifrage
- Saxifraga aizoides – Yellow mountain saxifrage,<ref name="Quattrocchi">Umberto Quattrocchi. CRC World Dictionary of Plant Names: Common Names, Scientific Names, Eponyms. Synonyms, and Etymology. CRC Press, 1999. p.2395-2396. Template:ISBN</ref> yellow saxifrage<ref name="Quattrocchi"/>
- Saxifraga aizoon – Aizoon rockfoil
- Saxifraga algisii
- Saxifraga anadyrensis
- Saxifraga androsacea
- Saxifraga aquatica
- Saxifraga arachnoidea
- Saxifraga × arendsii – mossy saxifrage, mossy rockfoil
- Saxifraga aspera L. – rough saxifrage,<ref name="Quattrocchi"/> stiff-haired saxifrage<ref name="Quattrocchi"/>
- Saxifraga bicuspidata
- Saxifraga biflora
- Saxifraga bronchialis L. – matte saxifrage
- Saxifraga bryoides L. – mossy saxifrage<ref name="Quattrocchi"/>
- Saxifraga burseriana L. – AGM
- Saxifraga caesia – blue green saxifrage<ref name="Quattrocchi"/>
- Saxifraga callosa Sm. – limestone saxifrage<ref name="Quattrocchi"/>
- Saxifraga canaliculata
- Saxifraga carpatica
- Saxifraga cernua – drooping saxifrage, nodding saxifrage, bulblet saxifrage
- Saxifraga cervicornis
- Saxifraga cespitosa – tufted saxifrage
- Saxifraga ciliata
- Saxifraga cochlearis – spoon-leaved saxifrage
- Saxifraga columnaris Schmalh.
- Saxifraga corsica
- Saxifraga consanguinea W.W.Sm.
- Saxifraga cotyledon L. – great alpine rockfoil, greater evergreen saxifrage
- Saxifraga crustata Vest. – crusted-leaved saxifraga, silver saxifrage, encrusted saxifrage
- Saxifraga cuneifolia – shield-leaved saxifrage,<ref name="Quattrocchi"/> lesser London pride
- Saxifraga cymbalaria – celandine saxifrage
- Saxifraga decipiens
- Saxifraga dinnikii Schmalh.
- Saxifraga eschscholtzii – cushion saxifrage
- Saxifraga exarata – furrowed saxifrage<ref name="Quattrocchi"/>
- Saxifraga flagellaris Willd. ex Sternb. – whiplash saxifrage, spider saxifrage, "spider plant"
- Saxifraga florulenta
- Saxifraga forbesei
- Saxifraga fortunei Hook.f. – fortune saxifrage
- Saxifraga × geum – Robertsoniana saxifrage (S. hirsuta x S. umbrosa)
- Saxifraga globulifera – Gibraltar saxifrage
- Saxifraga granulata L. – meadow saxifrage,<ref name="Quattrocchi"/> bulbous saxifrage,<ref name="Quattrocchi"/> fair maids of France<ref name="Quattrocchi"/> (type species)
- Saxifraga grisebachii - Engleria saxifrage
- Saxifraga groenlandica
- Saxifraga hederacea
- Saxifraga hirculus L. – yellow marsh saxifrage, marsh saxifrage, "bog saxifrage"
- Saxifraga hirsuta – kidney saxifrage<ref name="Quattrocchi"/>
- Saxifraga hyperborea – pygmy saxifrage
- Saxifraga hypnoides – mossy saxifrage,<ref name="Quattrocchi"/> Dovedale moss
- Saxifraga juniperifolia
- Saxifraga korshinskii Kom.
- Saxifraga lactea Turcz.
- Saxifraga longifolia – Pyrenean saxifrage<ref name="Quattrocchi"/>
- Saxifraga maderensis – Madeira saxifrage,<ref name="Quattrocchi"/> Madeira breakstone<ref name="Quattrocchi"/>
- Saxifraga mertensiana – Mertens' saxifrage
- Saxifraga montana
- Saxifraga moschata – musky saxifrage, mossy saxifrage
- Saxifraga moschata ssp. basaltica
- Saxifraga muscoides
- Saxifraga mutata
- Saxifraga nathorstii (Dusén) Hayek – East Greenland saxifrage
- Saxifraga nipponica
- Saxifraga oppositifolia – purple saxifrage,<ref name="Quattrocchi"/> purple mountain saxifrage
- Saxifraga osloensis Knaben – Oslo saxifrage, a natural hybrid species<ref>Template:Cite journal</ref>
- Saxifraga paniculata – lifelong saxifrage,<ref name="Quattrocchi"/> white mountain saxifrage
- Saxifraga paradoxa Sternb. – Fragile saxifraga
- Saxifraga petraea
- Saxifraga platysepala (= S. flagellaris auct. non Willd.) – broadsepal saxifrage
- Saxifraga porophylla
- Saxifraga redofskii – many-flower saxifrage
- Saxifraga rivularis – alpine brook saxifrage, brook saxifrage, highland saxifrage
- Saxifraga rosacea – Irish saxifrage
- Saxifraga rotundifolia L. – round-leaved saxifrage<ref name="Quattrocchi"/>
- Saxifraga roylei
- Saxifraga rudolphiana
- Saxifraga rufopilosa – redhair saxifrage
- Saxifraga sancta
- Saxifraga serpyllifolia – thymeleaf saxifrage
- Saxifraga sibirica – Siberian saxifrage
- Saxifraga spathularis – Saint Patrick's cabbage
- Saxifraga squarrosa – Dolomites saxifrage
- Saxifraga stolonifera – creeping saxifrage, strawberry saxifrage, creeping rockfoil, strawberry begonia, strawberry geranium,<ref name="Quattrocchi"/> Aaron's beard
- Saxifraga stolonifera f. aptera (Makino) H.Hara – hoshizaki-yukinoshita (Japanese)
- Saxifraga stolonifera 'Cuscutiformis' (Saxifraga cuscutiformis Lodd.) – Dodder-like saxifrage<ref name="Quattrocchi"/>
- Saxifraga subverticillata
- Saxifraga svalbardensis
- Saxifraga taygetea
- Saxifraga taylorii – Taylor's saxifrage
- Saxifraga tenella
- Saxifraga tombeanensis
- Saxifraga tricuspidata Rottb. – prickly saxifrage
- Saxifraga tridactylites – rue-leaved saxifrage, "nailwort"
- Saxifraga trifurcata
- Saxifraga × urbium – London pride (S. spathularis × S. umbrosa)
- Saxifraga umbrosa – Pyrenean saxifrage
- Saxifraga vandelli
- Saxifraga wahlenbergii
Formerly placed hereEdit
Plants formerly placed in Saxifraga are mainly but not exclusively Saxifragaceae. They include:Template:Citation needed
- Astilboides tabularis, as S. tabularis
- Bergenia crassifolia, as S. cordifolia, S. crassifolia
- Bergenia pacumbis, as S. ligulata, S. pacumbis
- Bergenia purpurascens, as S. delavayi, S. purpurascens
- Boykinia jamesii, as S. jamesii
- Boykinia occidentalis (Coastal Brookfoam), as S. elata
- Boykinia richardsonii (Richardson's Brookfoam), as S. richardsonii
- Darmera peltata (Indian Rhubarb), as S. peltata
- Leptarrhena pyrolifolia, as S. pyrolifolia
- Luetkea pectinata (Partridgefoot), as S. pectinata
- Micranthes, including:
- Micranthes integrifolia (wholeleaf saxifrage)
- Micranthes howellii (Howell's saxifrage),<ref name="Quattrocchi"/> as S. howellii
- Micranthes stellaris (Starry saxifrage),<ref name="Quattrocchi"/> as S. stellaris
- Mukdenia rossii (Mukdenia), as S. rossii
Other "saxifragous" plantsEdit
Several plant genera have names referring to saxifrages, although they might not be close relatives of Saxifraga. They include:Template:Citation needed
- Golden-saxifrages, Chrysosplenium
- Burnet-saxifrages, Pimpinella
- Pepper-saxifrage, Silaum silaus. The name "silaum" comes from the Latin word sil, which means yellow ochre. This refers to the sulphurous yellow colour of the flowers.<ref>Reader's Digest Nature Lover's Library Wild Flowers of Britain, page 192, published 1988</ref>
Some plants refer to Saxifraga in their generic names or specific epithets, either because they are also "rock-breaking" or because they resemble members of the saxifrage genus:Template:Citation needed
- Campanula saxifraga
- Celmisia saxifraga (Benth.) W.M.Curtis
- Cineraria saxifraga DC.
- Dryopteris saxifraga
- Petrorhagia saxifraga – Tunicflower
- Pimpinella saxifraga – Burnet saxifrage
- Ptychotis saxifraga
- Saxifragella
- Saxifragodes
- Saxifragopsis Small
EcologyEdit
Template:More citations needed section
Saxifrages are typical inhabitants of Arctic–alpine ecosystems, and are hardly ever found outside the temperate parts of the Northern Hemisphere; most members of this genus are found in subarctic climates. A good number of species grow in glacial habitats, such as S. biflora which can be found some Template:Cvt above sea level in the Alps, or the East Greenland saxifrage (S. nathorstii). The genus is also abundant in the Eastern and Western Himalayan alpine shrub and meadows. Though the archetypal saxifrage is a small plant huddling between rocks high up on a mountain, many species do not occur in such a habitat and are larger (though still rather delicate) plants found on wet meadows.
Various Saxifraga species are used as food plants by the caterpillars of some butterflies and moths, such as the Phoebus Apollo (Parnassius phoebus).<ref>Template:Cite book</ref>
Charles Darwin – erroneously believing Saxifraga to be allied to the sundew family (Droseraceae) – suspected the sticky-leaved round-leaved saxifrage (S. rotundifolia), rue-leaved saxifrage (S. tridactylites) and Pyrenean saxifrage (S. umbrosa) to be protocarnivorous plants, and conducted some experiments whose results supported his observations,<ref>Template:Cite book</ref> but the matter has apparently not been studied since his time.
CultivationEdit
Numerous species and cultivars of saxifrage are cultivated as ornamental garden plants, valued particularly as groundcover or as cushion plants in rock gardens and alpine gardens. Many require alkaline or neutral soil to thrive.<ref name=RHSAZ>Template:Cite book</ref>
S. × urbium (London pride), a hybrid between Pyrenean saxifrage (S. umbrosa) and St. Patrick's cabbage (S. spathularis), is commonly grown as an ornamental plant.<ref name="Cam"/> Another horticultural hybrid is Robertsoniana saxifrage (S. × geum), derived from kidney saxifrage (S. hirsuta) and Pyrenean saxifrage.Template:Citation needed Some wild species are also used in gardening. Cambridge University Botanic Garden hosts the United Kingdom's national collection of saxifrages.<ref name="Cam"/>
Award of Garden MeritEdit
The following species and cultivars have gained the Royal Horticultural Society's Award of Garden Merit:-<ref>{{#invoke:citation/CS1|citation |CitationClass=web }}</ref> Template:Div col
- 'Angelina Johnson' (fortunei)<ref>{{#invoke:citation/CS1|citation
|CitationClass=web }}</ref>
- 'Blackberry and Apple Pie' (fortunei)<ref>{{#invoke:citation/CS1|citation
|CitationClass=web }}</ref>
- S. callosa (limestone saxifrage)<ref>{{#invoke:citation/CS1|citation
|CitationClass=web }}</ref>
- 'Conwy Snow' (fortunei)<ref>{{#invoke:citation/CS1|citation
|CitationClass=web }}</ref>
- 'Coolock Kate'<ref>{{#invoke:citation/CS1|citation
|CitationClass=web }}</ref>
- 'Cumulus'<ref>{{#invoke:citation/CS1|citation
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- S. fortunei<ref>{{#invoke:citation/CS1|citation
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- 'Gregor Mendel' (× apiculata)<ref>{{#invoke:citation/CS1|citation
|CitationClass=web }}</ref>
- 'Lagraveana' (paniculata)<ref>{{#invoke:citation/CS1|citation
|CitationClass=web }}</ref>
- 'Lutea'<ref>{{#invoke:citation/CS1|citation
|CitationClass=web }}</ref>
- 'Minor'<ref>{{#invoke:citation/CS1|citation
|CitationClass=web }}</ref>
- 'Moe' (fortunei)<ref>{{#invoke:citation/CS1|citation
|CitationClass=web }}</ref>
- 'Monarch'<ref>{{#invoke:citation/CS1|citation
|CitationClass=web }}</ref>
- 'Mount Nachi' (fortunei)<ref>{{#invoke:citation/CS1|citation
|CitationClass=web }}</ref>
- 'Peach Melba'<ref>{{#invoke:citation/CS1|citation
|CitationClass=web }}</ref>
- 'Reginald Farrer' (Silver Farreri Group)<ref>{{#invoke:citation/CS1|citation
|CitationClass=web }}</ref>
- 'Rokujo' (fortunei)<ref>{{#invoke:citation/CS1|citation
|CitationClass=web }}</ref>
- 'Rosea'<ref>{{#invoke:citation/CS1|citation
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- 'Shiranami' (fortunei)<ref>{{#invoke:citation/CS1|citation
|CitationClass=web }}</ref>
- 'Slack's Ruby Southside' (Southside Seedling Group)<ref>{{#invoke:citation/CS1|citation
|CitationClass=web }}</ref>
- 'Snowflake' (Silver Farreri Group)<ref>{{#invoke:citation/CS1|citation
|CitationClass=web }}</ref>
- 'Southside Star' (Southside Seedling Group)<ref>{{#invoke:citation/CS1|citation
|CitationClass=web }}</ref>
- S. stolonifera (strawberry saxifrage)<ref>{{#invoke:citation/CS1|citation
|CitationClass=web }}</ref>
- 'Sue Drew' (fortunei)<ref>{{#invoke:citation/CS1|citation
|CitationClass=web }}</ref>
- Template:Tdes='Toujya' (fortunei)<ref>{{#invoke:citation/CS1|citation
|CitationClass=web }}</ref>
- 'Theoden'<ref>{{#invoke:citation/CS1|citation
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- 'Tumbling Waters' <ref>{{#invoke:citation/CS1|citation
|CitationClass=web }}</ref>
- S. × urbium (London pride)<ref>{{#invoke:citation/CS1|citation
|CitationClass=web }}</ref>
- 'Venetia' (paniculata)<ref>{{#invoke:citation/CS1|citation
|CitationClass=web }}</ref>
- 'Whitehill'<ref>{{#invoke:citation/CS1|citation
|CitationClass=web }}</ref> Template:Div col end
UsesEdit
The leaves of some saxifrage species, such as creeping saxifrage (S. stolonifera) and S. pensylvanica,<ref>Template:Cite book</ref> are edible. The former is a food in Korea<ref>Template:Cite journal</ref> and Japan.Template:Citation needed The flowers of purple saxifrage (S. oppositifolia) are eaten in Nunavut, Canada and the leaves and stems brewed as a tea.<ref name="nunavut">Official Flower of Nunavut, Nunavut, Canada</ref>
Species are also used in traditional medicine, such as creeping saxifrage in East Asia<ref>Template:Cite book</ref> and round-leaved saxifrage (S. rotundifolia) in Europe.<ref>Template:Cite book</ref>
Two species—purple saxifrage and creeping saxifrage—are popular floral emblems. They are official flowers for:
- Nunavut, Canada - purple saxifrage<ref>{{#invoke:citation/CS1|citation
|CitationClass=web }}</ref>
- County Londonderry, Northern Ireland - purple saxifrage<ref>County flowers in Britain Template:Webarchive www.plantlife.org.uk</ref>
- Tsukuba, Japan - creeping saxifrage, "hoshizaki" form (S. stolonifera Curtis f. aptera)<ref>{{#invoke:citation/CS1|citation
|CitationClass=web }}</ref>
- Saxifraga umbrosa a3.jpg
Pyrenean saxifrage (S. umbrosa), ancestor to horticultural hybrid saxifrages
- Saxifraga caesia a1.jpg
- Saxifraga cuneifolia3.jpg
Lesser London pride (Saxifraga cuneifolia)
- Saxifraga decipiens White V08 H3990.jpg
- Saxifraga rosacea 01.jpg
Irish saxifrage (Saxifraga rosacea)
- Saxifraga tricuspidata upernavik 2007-07-15 1 filtered.jpg
Prickly saxifrage (Saxifraga tricuspidata) flowers
ReferencesEdit
BibliographyEdit
- Books
- Articles
- Template:Cite EB1911
- Template:Cite journal
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- Websites
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- Floras
- {{#invoke:citation/CS1|citation
|CitationClass=web }}, in Flora of China online vol. 8 see also PDF
- {{#invoke:citation/CS1|citation
|CitationClass=web }} Template:Refend