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The vireos Template:IPAc-en make up a family, Vireonidae, of small to medium-sized passerine birds found in the New World (Canada to Argentina, including Bermuda and the West Indies) and Southeast Asia. The family contains 62 species and is divided into eight genera. "Vireo" is a Latin word referring to a green migratory bird, perhaps the female golden oriole, possibly the European greenfinch.<ref name=job>Template:Cite book</ref><ref name=OED>Template:Cite OED</ref>

They are typically dull-plumaged and greenish in color, the smaller species resembling wood warblers apart from their heavier bills. They range in size from the Chocó vireo, dwarf vireo and lesser greenlet, all at around 10 cm and 8g, to the peppershrikes and shrike-vireos at up to 17 cm and 40g.<ref name=for>Forshaw, Joseph & Parkes, Kenneth C. 1991. Encyclopaedia of Animals: Birds, p. 209. Merehurst Press, London. Template:ISBN</ref>

Distribution and habitatEdit

Most species are found in Middle America and northern South America. Thirteen species of true vireos occur farther north, in the United States, Bermuda<ref>White-eyed vireo, Audubon Society of Bermuda</ref> and Canada; of these all but Hutton's vireo are migratory. Members of the family seldom fly long distances except in migration.<ref name="sal">Salaman, Paul & Barlow, Jon C. 2003. Vireos. Pp. 478–479 in; Perrins, C. ed. The Firefly Encyclopedia of Birds. Firefly Books, Oxford. Template:ISBN</ref> They inhabit forest environments, with different species preferring forest canopies, undergrowth, or mangrove swamps.<ref name=for/>

A few species in the genus Vireo have appeared on the eastern side of the Atlantic as vagrants to the Western Palearctic.<ref>{{#invoke:citation/CS1|citation |CitationClass=web }}</ref>

BehaviourEdit

The resident species occur in pairs or family groups that maintain territories all year (except Hutton's vireo, which joins mixed feeding flocks). Most of the migrants defend winter territories against conspecifics. The exceptions are the complex comprising the red-eyed vireo, the yellow-green vireo, the black-whiskered vireo, and the Yucatan vireo, which winter in small wandering flocks.<ref name="sal"/>

VoiceEdit

File:Cyclarhis gujanensis.jpg
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Males of most species are persistent singers. Songs are usually rather simple, monotonous in some species of the Caribbean littoral and islands, and most elaborate and pleasant to human ears in the Chocó vireo and the peppershrikes.<ref name="sal"/>

BreedingEdit

The nests of many tropical species are unknown. Of those that are known, all build a cup-shaped nest that hangs from branches. The female does most of the incubation, spelled by the male except in the red-eyed vireo complex.<ref name="sal"/>

FeedingEdit

All members of the family eat some fruit but mostly insects and other arthropods. They take prey from leaves and branches; true vireos also flycatch, and the gray vireo takes 5 percent of its prey from the ground.<ref name="sal"/>

SystematicsEdit

The family Vireonidae is related to the crow-like birds in family Corvidae and the shrikes in family Laniidae as part of superfamily Corvoidea. As currently circumscribed the family is made up of eight genera.<ref name=IOC>Template:Cite taxon</ref><ref name=BOW>Template:Cite BOW</ref>

Traditionally the family was considered to include four New World genera containing the true vireos (Vireo), the greenlets (Hylophilus), the shrike-vireos (Vireolanius) and the peppershrikes (Cyclarhis). However, phylogenetic studies found Hylophilus to be polyphyletic, with the greenlets split into three distinct groups: the "scrub" greenlets in a restricted Hylophilus, the "canopy" greenlets in resurrected genus Pachysylvia and the tawny-crowned greenlet in new genus Tunchiornis.<ref>Template:Cite journal</ref><ref>Template:Cite journal</ref><ref>{{#invoke:citation/CS1|citation |CitationClass=web }}</ref>

In addition, biochemical studies have identified two babbler genera (Pteruthius and Erpornis) which may be Old World members of this family.<ref name=reddy>Reddy, Sushma & Cracraft, Joel (2007): Old World Shrike-babblers (Pteruthius) belong with New World Vireos (Vireonidae). Molecular Phylogenetics and Evolution, 44 (3): 1352–1357. {{#invoke:doi|main}}</ref> Observers have commented on the vireo-like behaviour of the Pteruthius shrike-babblers, but apparently no-one suspected the biogeographically unlikely possibility of vireo relatives in Asia. Some recent taxonomic treatements, such as the IOC taxonomy followed here, include Pteruthius and Erpornis in Vireionidae,<ref name=IOC/><ref name=BOW/> whereas other place them in their own families Pteruthidae and Erpornidae.<ref>Template:Cite journal</ref>

Species in taxonomic orderEdit

Image Genus Species
File:Black-eared Shrike Babbler Khangchendzonga national Park West Sikkim India 30.10.2015.jpg Pteruthius - shrike-babblers<ref name=reddy/> Template:Small
File:Erpornis zantholeuca.jpg Erpornis <ref>Barker, F. Keith; Cibois, Alice; Schikler, Peter A.; Feinstein, Julie & Cracraft, Joel (2004): "Phylogeny and diversification of the largest avian radiation." PNAS 101(30): 11040-11045. {{#invoke:doi|main}} Template:PMID Supporting information</ref> Template:Small
File:Pitiguari (Cyclarhis gujanensis) (17359710466).jpg Cyclarhis Template:Small, the peppershrikes
File:Vireolanius pulchellus -Panama-8a 2.jpg Vireolanius Template:Small, the shrike-vireos
File:Hylophilus semicinereus - Grey-chested Greenlet.JPG Hylophilus Template:Small
File:Tawny-crowned Greenlet (Hylophilus ochraceiceps) (7222935148).jpg Tunchiornis Template:Small
File:Dusky-capped Greenlet (Hylophilus hypoxanthus).jpg Pachysylvia Template:Small
File:Edit white-eyed vireo blandair 4.20.20 DSC 0224.jpg Vireo Template:Small, the true vireos.

ReferencesEdit

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External linksEdit

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